| 0. Squamose vestiture of the epistome: covered with scales similar in size, density and color to those of remainder of rostrum (0) (Fig. 4D); covered with scales smaller, sparser and usually of different color to those of remainder of rostrum (1) (Fig. 5E); lacking scales (2). |
| 1. Vestiture of rostrum: setae on rostrum not forming semi-circles on sides of rostral sulcus (0); forming semi circles on sides of rostral sulcus (1) (Fig. 5C). |
| 2. Vestiture of antennae: scape and funicle scarcely setose (0); scape squamose and funicle setose (1); scape and funicle squamose (2). |
| 3. Vestiture, verticilate setae of antennal funicle (coarse, stiff setae around apex of antennal articles): setae light-colored (0); setae dark-colored (1). |
| 4. Vestiture of scutellum: vestiture present, consisted on scales similar in color to those covering whole surface of elytra (0); vestiture present, consisted on white scales or seta-like scales (1) (Fig. 2B); vestiture absent (2). |
| 5. Vestiture, dark maculae of seta-like scales on pronotum and elytra: dark maculae absent 0); dark maculae present (1) (Fig. 1E). |
| 6. Scaly vestiture of elytra: uniformly present on whole surface of elytra (0) (Fig. 1B); forming longitudinal stripes or maculae and leaving glabrous areas (1) (Fig. 2A); elytra covered with seta-like scales (2) (Fig. 2E); elytra lacking scaly vestiture (3) (Fig. 2D). |
| 7. Setose vestiture of elytra: absent (0) (Fig. 2D); setae short, recumbent to suberect, either fine or scale-like (1) (Fig. 3F); setae erect, long (about as long as width of interstria 2 near middle) (2) (Fig. 2E); setae erect, very long (longer than width of elytral interstriae 2 near middle) (3) (Fig. 3B). |
| 8. Rostrum, expansion of apex in dorsal view (including borders of scrobes): rostrum expanded at apex, wider than frons (0) (Fig. 4A); rostrum not expanded at apex, equal to narrower than frons (1) (Fig. 5B). |
| 9. Rostrum, curvature of dorso-lateral margins: dorso-lateral margins divergent towards apex (0) (Fig. 4B); margins subparallel to convergent towards apex (1) (Fig. 5B); margins curved and convergent towards apex in males and subparallel in females (2); margins curved and convergent towards apex in both sexes (3) (Fig. 4E). |
| 10. Rostrum, curvature in profile view: rostrum with dorsal surface convex (0); rostrum with flat or depressed dorsal surface (1). |
| 11. Rostrum, pair of lateral carinae: carinae indistinct (0) (Fig. 4F); carinae subparallel to convergent towards frons (1) (Fig. 5A); carinae divergent towards frons (2) (Fig. 5C). |
| 12. Rostrum, borders of scrobes: not to slightly thickened (0) (Fig. 5F); strongly thickened (1) (Fig. 4B). |
| 13. Rostrum, apex of scrobes in dorsal view: scrobes completely exposed from above (0) (Fig. 4A); scrobes slightly visible from above (1) (Fig. 4C). |
| 14. Rostrum, scrobes in lateral view: slightly curved, posteriorly shallow and oriented towards eye (0); variously curved downwards towards ventral margin of eye (1) (Fig. 2E); strongly curved downward (almost 90°) and directed ventrad of head (2) (Fig. 3E). |
| 15. Rostrum, elevation of epistome: epistome not elevated relative to post-rostrum (0) (Fig. 5B); epistome elevated relative to post-rostrum (1) (Fig. 5D). |
| 16. Rostrum, nasal plate sensu Vaurie, 1963: large, triangular, not descending, separated from epistome by a carina or elevated border (0) (Fig. 4B); narrow, V-shaped, descending, not separated from epistome by a carina or elevation (1) (Fig. 5A); very narrow, descending, not visible from above (2) (Fig. 4E). |
| 17. Rostrum, presence of median groove: median groove absent along rostrum (0) (Fig. 4A); median groove always present along rostrum and frons (1) (Fig. 5E). |
| 18. Rostrum, ventral side, position and extension of occipital sutures: occipital sutures well-developed, transversal (0); occipital sutures well-developed, oblique (1); occipital sutures reduce to indistinct (2) [See Franz, 2012, Fig. 30, char. 25.0, and Fig. 3, char. 25.1]. |
| 19. Rostrum, ventral side, hypostomal-labial sutures: ventral side without large triangular impression flanked by foveate hypostomal-labial sutures (0); with elongate triangular impression flanked by long, non-foveate hypostomal-labial sutures (1) [See Franz, 2012, Fig. 28, char. 24(0) and Fig. 29, char. 24(1)]. |
| 20. Mouthparts, setation of prementum: prementum lacking setae (0); prementum with various long setae (1). |
| 21. Head, pair of arched grooves on sides of frons, around eyes: absent (0); present (1) (Fig. 5F). |
| 22. Head, presence of preocular sulcus: preocular sulcus present (0) (Fig. 4C); preocular sulcus absent (1) (Fig. 4D). |
| 23. Head, supraocular borders: not thickened (0); thickened (1) (Fig. 1F). |
| 24. Head, position of eyes: eyes distinctly encroaching frons (0) (Fig. 4B); slightly encroaching frons (1) (Fig. 4D); entirely lateral (2) (Fig. 4F). |
| 25. Head, shape of eyes: eyes oval, dorso-ventrally oriented (0) (Fig. 3E); eyes rounded to slightly oval and antero-posteriorly oriented (1) (Fig. 1E). |
| 26. Head, postocular constriction: constriction slight to absent (0), constriction strong (1) (Fig. 5B). |
| 27. Head, lateral view: gular angle very obtuse (0); slightly obtuse (1); almost right angled (2). |
| 28. Antennae, cross-sectional shape of scape: scape cylindrical in cross section (0) (Fig. 5A); scape flat in cross section (1) (Fig. 4E). |
| 29. Antennae, shape of scape: scape clavate (0) (Fig. 4D); scape capitate (1) (Fig. 5C); scape strongly clavate (2) (Fig. 4F); scape spatulate (3) (Fig. 4E). |
| 30. Antennae, curvature of scape: scape not curved (0) (Fig. 5A); scape curved (1) (Fig. 5D). |
| 31. Antennae, length of scape with respect to hind margin of eye or apex of pronotum: scape not reaching hind margin of eye (0) (Fig. 4C); reaching to slightly exceeding hind margin of eye (1) (Fig. 5D); reaching to slightly exceeding apex of pronotum (2) (Fig. 2F); largely exceeding apex of pronotum (3) (Fig. 4A). |
| 32. Antennae, sexual dimorphism: scape width subequal in males and females (0); scape very broad in males and slender in females (1). |
| 33. Antennae, length of funicular article 2 (antennomere 3) with respect to length of funicular article 1 (antennomere 2): funicular article 2 slighlty shorter to about as long as article 1 (0); funicular article 2 less than 2× longer than article 1 (1); funicular article 2, more than 2× longer than article 1 (2). |
| 34. Antennae, length to width ratio of funicular article 3 (antennomere 4): funicular article 3 about as long as, to slightly longer than wide (0); distinctly elongate (more than 2× as long as wide) (1). |
| 35. Antennae, length to width ratio of funicular article 7 (antennomere 8): funicular article 7 wider than long or as long as wide at apex (0); distinctly longer than wide at apex (1). |
| 36. Antennae, length of funicular article 3 (antennomere 4) regarding funicular article 7 (antennomere 8): funicular article 3 longer than funicular article 7 (0); funicular article 3 as long as, to shorter than article 7 (1). |
| 37. Antennae, shape of club: club oval, very short (length/width 2 to 2.25×) (0); club oval, moderately short (length/width 2.26 to 3×) (1); fusiform, very long (length/width more than 3×) (2). |
| 38. Pronotum, shape in dorsal view: subcylindrical (sides curved, maximum width about middle) (0) (Fig. 2D); slightly conical (sides divergent towards middle and subparallel on posterior half) (1) (Fig. 3B); strongly conical (sides strongly divergent from apex to base) (2) (Fig. 1B). |
| 39. Pronotum, vibrissae on anterior margin: vibrissae absent (0); vibrissae present (1). |
| 40. Pronotum, lateral tubercles at about middle of flanks: tubercles absent (0); tubercles present (1) (Fig. 1C). |
| 41. Pronotum, median impression, longitudinal groove and/or elevation: disc of pronotum with fine, usually interrupted median groove (0); disc of pronotum with broad median groove (1) (Fig. 1C); disc of pronotum with median impression and groove (2) (Fig. 1A); disc of pronotum elevated along midline (3). |
| 42. Pronotum, pair of lateral, longitudinal impressions: lateral impressions absent (0); lateral impressions present (1) (Fig. 1A). |
| 43. Pronotum, pairs of short, oblique lateral sulci: lateral sulci absent (0); lateral sulci present (1) (Fig. 3A). |
| 44. Pronotum, curvature of disc in lateral view: flat to slightly convex (0); strongly convex (1) (Fig. 2F). |
| 45. Pronotum, postero-lateral angles (females): postero-lateral angles not projecting (0) (Fig. 2C); postero-lateral angles projecting (1); postero-lateral angles projecting in a tubercle (2) (Fig. 2B). |
| 46. Pronotum, constriction and thickness of margins: anterior and posterior margins not constricted, not to slightly thickened (0); posterior margin constricted and thickened (1) (Fig. 2B); anterior and posterior margins strongly constricted and thickened (2) (Fig. 3D). |
| 47. Pronotum, outline of base: base straight to slightly bisinuate (0); base strongly bisinuate (1) (Fig. 3A). |
| 48. Pronotum, sculpture in dorsolateral profile: disc of pronotum smooth to slightly punctuate (0) (Fig. 2D); disc of pronotum with irregular foveae or large punctures (1) (Fig. 3A); disc of pronotum granulose (2) (Fig. 3C); disc of pronotum striate (3). |
| 49. Scutellum, shape: scutellum quadrate to rounded (0); scutellum triangular (1); scutellum reduce to indistinct (2). |
| 50. Elytra, outline of base: base of elytra straight (0) (Fig. 2C); base of elytra slightly bisinuate (1) (Fig. 1C); base of elytra strongly bisinuate (2) (Fig. 3A). |
| 51. Elytra, development of humeri: humeri well-developed (0) (Fig. 2A); humeri reduced (1) (Fig. 3C); humeri absent (2) (Fig. 2C). |
| 52. Elytra, presence of humeral tubercle: humeri lacking tubercle (0); humeri projecting in a small anteriorly directed tubercle (1) (Fig. 1D); humeri projecting in a postero-laterally directed tubercle (2) (Fig. 1C); humeri projecting in a long laterally directed tubercle (3) (Fig. 2D). |
| 53. Elytra, presence of hump: elytra not humped (not ascending towards declivity) (0) (Fig. 2E); elytral disc humped (ascending towards declivity) (1) (Fig. 1E). |
| 54. Elytra, height in lateral view: elytra very high (0) (Fig. 3E); elytra moderately high (1) (Fig. 2E); elytra flat (2) (Fig. 2F). |
| 55. Elytra, development of subapical calli: subapical calli well-developed (0) (Fig. 3A); subapical calli slightly developed (Fig. 2B) (1); subapical calli indistinct (2) (Fig. 2C). |
| 56. Elytra, apical declivity: apical declivity abrupt (0) (Fig. 3F); apical declivity slight to moderate (1) (Fig. 2F). |
| 57. Elytra, presence of tubercles on disc and declivity: elytra lacking tubercles (0); elytra with tubercles (1) (Fig. 1F). |
| 58. Elytra, outline of apex: apex entire (0); apex bifid (1) (Fig. 3D). |
| 59. Elytra, curvature of intervals (= interstriae): intervals flat (0) (Fig. 1B); intervals slightly convex (1); uneven intervals (except suture) strongly convex, even intervals flat (2) (Fig. 1A); all intervals strongly convex (3) (Fig. 2D). |
| 60. Elytra, presence of supernumerary striae: supernumerary striae absent (0); supernumerary striae (more than 10 striae in each elytrum) present (1) (Fig. 1A). |
| 61. Elytra, proximity of striae 9º and 10º: elytra with striae 9º and 10º fused along posterior ⅔ (0); slightly closer to each other along posterior ⅔ (1); well separated along their extension (2). |
| 62. Elytra, size of punctures of striae: punctures large (about as wide as, to slightly wider than intervals) (0) (Fig. 3A); punctures medium-sized to small (narrower than intervals) (1) (Fig. 1A); punctures very small and close to each other (2) (Fig. 1B). (The ratio width of interval/width of puncture was measured on intervals 2 to 5, about middle length of elytra). |
| 63. Metathoracic wings, development: wings well-developed (0); wings reduced to absent (1). |
| 64. Legs, separation of front coxae (females): front coxae contiguous (0); front coxae separated from each other (1). |
| 65. Legs, distance of front coxae regarding margins of prosternum: front coxae closer to posterior than to anterior margin of prosternum (0); equally separated from anterior and posterior margins of prosternum or slightly closer to anterior margin (1); twice as close to anterior margin than to posterior margin of prosternum (2). |
| 66. Legs, width of front femora relative to width of hind tibiae (females): hind femora wider than front femora (0) (Fig. 3E); front femora about as wide as, to slightly wider (up to 1.25×) than hind femora (1) (Fig. 1E); front femora wider than hind femora (more than 1.25× and less than 1.50×) (2); front femora strongly widened (1.50× or more, as wide as hind femora) (3) (Fig. 3D). |
| 67. Legs, presence of denticles in front femora: denticles of femora absent (0); front femora with one to three small denticles on inner side (1); all femora with one large denticle (2). |
| 68. Legs, size of mucro of front tibiae: mucro indistinct (0); mucro small to medium-sized (less than half, or about as long as half width of tibial apex) (1); mucro large (about as long as width of tibial apex) (2). |
| 69. Legs, front tibiae: without prominence opposite to mucro (0); with prominence opposite to mucro (1). |
| 70. Legs, presence of row of denticles on inner margin of tibiae: denticles absent to minute (0); denticles present in front tibiae only (1); denticles present in the three pairs of tibiae (2). |
| 71. Legs, corbel at metatibial apex: corbel broad, squamose (2 to 2.5× as long as wide) (0); corbel narrow squamose or setose (more than 3× as long as wide) (1); corbel lacking (= metatibial apex simple) (2). |
| 72. Legs, length of dorsal comb relative to length of distal comb of hind tibiae: dorsal comb shorter than distal comb (0); dorsal comb about as long as distal comb (1); dorsal comb longer than dorsal comb (2). [See Buchanan, 1939, pag. 7, Figs. 2J, 2B, 2I]. |
| 73. Legs, length relative to width of tarsite 2 of hind tibiae: tarsite 2 wider than long (0); tarsite 2 about as long as wide (1); tarsite 2 longer than wide (2). |
| 74. Venter, width of intercoxal portion of first abdominal ventrite: intercoxal portion about as wide as, to slightly narrower than space of hind coxae (0); intercoxal portion distinctly wider than space of hind coxae (1). |
| 75. Venter, presence of impression on ventrites 1 and 2 (females): ventrites 1 and 2 flat, lacking impression (0); ventrites 1 and 2 with a median impression (1). |
| 76. Venter, length of ventrite 2 relative to length of ventrites 3 + 4, measured along midline: ventrite 2 twice as long as ventrites 3 + 4 (0); slightly longer than ventrites 3 + 4 (1); about as long as ventrites 3 + 4, to slightly shorter (2). |
| 77. Venter, length of ventrite 1 regarding relative to ventrite 2, measured along midline: ventrite 1 distinctly longer than ventrite 2 (0); ventrite 1 about as long as ventrite 2 (1). |
| 78. Female terminalia, shape of sternite VIII: subtriangular (0) (Fig. 6A); subrhomboidal, very elongate (1) (Fig. 6B); subrhomboidal, slightly elongate (2) (Fig. 6C); rounded to suboval (3) (Fig. 6D); subcordiform (4) (Fig. 6E); subpentagonal (= shovel-shaped) (5) (Fig. 6F). |
| 79. Female terminalia, apical margin of sternite VIII: sternite VIII entire (0); sternite VIII divided (1); sternite VIII divided and projected in a pair of denticles (2) (Fig. 6G). |
| 80. Female terminalia, length of apodeme of sternite VIII: apodeme less than 2× as long as plate (0) (Fig. 6A); apodeme equally long or more than 2× as long as plate (1) (Fig. 6D). |
| 81. Female terminalia, length of ovipositor (distal plus dorsal gonocoxites): ovipositor about ⅓ of abdominal length (0); ovipositor about half length of abdomen (1); ovipositor about – of abdominal length (2); about as long as, to slightly longer than abdomen (3). |
| 82. Female terminalia, presence of baculi along ovipositor: baculi absent (0) (Fig. 7A); baculi present (1) (Fig. 7B). |
| 83. Female terminalia, presence of rows of setae along each side of baculi: ovipositor lacking rows of setae (0); ovipositor with rows of setae (1) (Fig. 7B). |
| 84. Female terminalia, shape and degree of sclerotization of distal gonocoxites: distal gonocoxites slightly to moderately sclerotized, not projected (0) (Fig. 7B); strongly sclerotized and projected in a short nail-shaped piece (1) (Fig. 7C); strongly sclerotized and strongly projected in a long nail- shaped piece (2) (Fig. 7D). |
| 85. Female terminalia, presence of styli: styli present, visible from ventral side (0) (Fig. 7B); styli present, hidden by distal gonocoxites from ventral side (1) (Fig. 7E); styli absent (2) (Fig. 7C). |
| 86. Spermatheca, width of spermathecal duct: spermathecal duct fine (0) (Fig. 8D); spermathecal duct wide (1) (Fig. 8B). |
| 87. Spermatheca, undulation of spermathecal duct: spermathecal duct straight to slightly undulate (Fig. 8A) (0); spermathecal duct curled (1) (Fig. 8B). |
| 88. Spermatheca, sclerotization of spermathecal duct: spermathecal duct slightly sclerotized (0); spermathecal duct as sclerotized as spermatheca (1). |
| 89. Spermatheca, shape of corpus: corpus subcylindrical, slender (about as wide as base of cornu) (0) (Fig. 8B); corpus broadly subcylindrical (1) (Fig. 8C); corpus amphora-shaped (2) (Fig. 8D); corpus subspherical (3) (Fig. 8F). |
| 90. Spermatheca, position of ramus: ramus inserted laterally on corpus of spermatheca (0) (Fig. 8G); ramus inserted on proximal extreme of corpus, same as collum (1) (Fig. 8E). |
| 91. Spermatheca, development of ramus: ramus distinct (0) (Fig. 8G); ramus indistinct (1) (Fig. 8A). |
| 92. Spermatheca, shape and length of collum (= nodulus): collum indistinct (0) (Fig. 8D); conical, short (1) (Fig. 8A); collum conical, moderately long (2) (Fig. 8B); collum subcylindrical, long, curved in opposite direction regarding apex of cornu (3) (Fig. 8G); collum subcylindrical, long, extended in same direction of corpus (4) (Fig. 8H). |
| 93. Spermatheca, basal prominence of collum: basal prominence absent (0); basal prominence present (1) (Fig. 8B). |
| 94. Spermatheca, length of cornu: cornu long (longer than corpus) (0) (Fig. 8G); medium length (about as long as corpus) (1) (Fig. 8C); short (shorter than corpus) (2) (Fig. 8D). |
| 95. Aedeagus, length of apodemes (= temones): apodemes about as long as median lobe (0) (Fig. 9G); apodemes about half length of median lobe (1) (Fig. 9F); apodemes twice as long as median lobe (2). |
| 96. Aedeagus, shape of apex of median lobe: apex acute to slightly acute (0) (Fig. 9A); apex truncate (1) (Fig. 9B); apex rounded (Ericydeus type) (2) (Fig. 9C); apex arrow- shaped (3) (Fig. 9D); apex horn- shaped (4) (Fig. 9E). |
| 97. Aedeagus, presence of setae on apex: setae absent (0) (Fig. 9F); setae present around ostium (1) (Fig. 9G); setae present around ostium and on ventral surface (2) (Fig. 9H). |
| 98. Aedeagus, presence of flagellum: flagellum absent (0); flagellum present (1). |
| 99. Aedeagus, sclerites of the internal sac: sclerites absent or not Cyrtomon type (0); sclerites consisted on a pair of lateral struts on each side of a pyriform piece connected with ejaculatory duct = Cyrtomon type (1) (Fig. 9I). |
