Two new records of terrestrial isopods (Isopoda, Oniscidea) from the state of Paraíba, Brazil

buckupi Campos-Filho & Cardoso, 2018 and Atlantoscia floridana (Van Name, 1940) have their distribution extended, and a short discussion about the record of A. buckupi in a semiarid area is provided.

In this work two species of terrestrial isopods are recorded from the state of Paraíba for the first time: Cubaris murina Brandt, 1833 from Campina Grande and Cabaceiras, and Porcellionidaes pruinosus (Brandt, 1833) from Campina Grande. In addition, A. buckupi and A. floridana were also found to occur in Campina Grande, the former in the Caatinga biome, which will be briefly discussed below.

MaTerial and MeThods
The material was collected with the aid of pitfall traps as proposed by Aquino et al. (2006). The traps consisted of Styrofoam containers 10 cm high and 10 cm diameter with a solution of 100 ml ethanol 70% and 5 ml of detergent. The sampling was carried out during the dry season, from 9 to 13 October 2017. The soil salinity and pH levels of the sampling site were measured with aid of TDS&EC B-M1X and ATC 0-14 pen type equipment, respectively. The material was stored in 75% ethanol and identifications were based on morphological characters. The species were examined with the aid of Biofocus SQFL-BI and Olympus CH-2 microscopes and, when necessary, appendages were mounted in micro-preparations with Hoyer's medium (Anderson, 1954). The material is deposited in the Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil (MZUSP).

Study area
The state of Paraíba is located in northeastern Brazil with an area of about 56,000 km 2 (3.6% of the Northeastern Region total area). The state is mainly covered by Caatinga and Atlantic forest (MMA, 2015;SUDENE, 2017). The Caatinga formation has a large territorial coverage, more than 900,000 km 2 (approximately 54% of the territory of the Northeastern Region and 11% of the whole Brazilian territory), and it composes the vegetation mosaic of the Seasonal Tropical Dry forests (Andrade-Lima, 1981;Silva et al., 2011;Pereira et al., 2017;SUDENE, 2017). According to Köppen's criteria, the state has three distinct climate regions: 1) littoral, ' Agreste' and 'Brejos de Altitude' with rainfall of about 1,800 mm/year; 2) 'Sertão' , with rainfall of about 820 mm/year; and 3) 'Borborema Plateau' , with rainfall of about 800 mm/year (Francisco et al., 2015).

disCussion
To date, the Caatinga comprises a high biodiversity with a high number of endemics (Leal et al., 2005;Loebmann & Haddad, 2010), and, as other Brazilian biomes, suffers with high levels of degradation due to overexploitation of its natural resources (Alves et al., 2009).
Among the species mentioned here, the most intriguing is the presence of A. buckupi in a semiarid area. To date, all species of Alloniscus are distributed in coastal areas, except the doubtful A. simplex Schmölzer, 1974 from Aberdare mountains, Kenya (Schmalfuss, 2003;Campos-Filho et al., 2018b). The specimens of A. buckupi were collected near an artificial lake within UFCG (Fig. 1A), where the dissolved salt and pH levels strongly exceed sea levels (1,282 ppm and 10.8 vs. 35 ppm and 7.4-8.5). Salt dependency related with physiology have been published in different groups of littoral species (e.g., Moens and Vincx, 2000;Braby and Somero, 2009;Pétillon et al., 2011;Sandman et al., 2013). Within Oniscidea, many papers regarding the systematics of coastal species have been published (e.g., Taiti and Howarth, 1996;Taiti et al., 2003;2018;Taiti and Lopez, 2008;Messina et al., 2011;2012;Lisboa el al., 2017;Pérez-Schultheiss et al., 2018), but only a few studies approached the relationships between species and salt environments (e.g., Barnes, 1932;1934;1935;Zimmer et al., 2002;Lopes-Leitzke et al., 2009;. Alloniscus buckupi is considered to be introduced to Campina Grande and, probably, the salt levels of the artificial lake allow the species to survive in this area. All species of the genus are diggers on sandy beaches, probably to avoid water loss and predation (see Schmalfuss, 1884). Campos-Filho et al. (2018b) mentioned that specimens of A. buckupi were collected during a drizzle in Cabo Branco beach, João Pessoa. The specimens from Campina Grande were collected from soil samples, confirming the digger habit of this species. referenCes