ISSN 0031-1049 printed version
ISSN 1807-0205 online version



General Information

Papéis Avulsos de Zoologia (PAZ) and Arquivos de Zoologia (AZ) cover primarily the fields of Zoology, publishing original contributions in systematics, paleontology, evolutionary biology, ontogeny, faunistic studies, and biogeography. Papéis Avulsos de Zoologia and Arquivos de Zoologia also encourage submission of theoretical and empirical studies that explore principles and methods of systematics.

All contributions must follow the International Code of Zoological Nomenclature. Relevant specimens should be properly curated and deposited in a recognized public or private, non-profit institution. Tissue samples should be referred to their voucher specimens and all nucleotide sequence data (aligned as well as unaligned) should be submitted to GenBank ( or EMBL (


Peer Review

All submissions to Papéis Avulsos de Zoologia and Arquivos de Zoologia are subject to review by at least two referees and the Editor-in-Chief. All authors will be notified of submission date. Authors may suggest potential reviewers. Communications regarding acceptance or rejection of manuscripts are made through electronic correspondence with the first or corresponding author only. Once a manuscript is accepted providing changes suggested by the referees, the author is requested to return a revised version incorporating those changes (or a detailed explanation of why reviewer's suggestions were not followed) within fifteen days upon receiving the communication by the editor.



Page-proofs with the revised version will be sent to e mail the first or corresponding author. Page-proofs must be returned to the editor, preferentially within 48 hours. Failure to return the proof promptly may be interpreted as approval with no changes and/or may delay publication. Only necessary corrections in proof will be permitted. Once page proof is sent to the author, further alterations and/or significant additions of text are permitted only at the author's expense or in the form of a brief appendix (note added in proof).


Submission of Manuscripts

Manuscripts should be sent to the SciELO Submission, along with a submission letter explaining the importance and originality of the study. Address and e mail of the corresponding author must be always updated since it will be used to send the 50 reprints in titled by the authors. Figures, tables and graphics should not be inserted in the text. Figures and graphics should be sent in separate files with the following formats: ".jpg" and ".tif" for figures, and ".xls" and ".cdr" for graphics, with 300 dpi of minimum resolution. Tables should be placed at the end of the manuscript.

Manuscripts are considered on the understanding that they have not been published or will not appear elsewhere in substantially the same or abbreviated form. The criteria for acceptance of articles are: quality and relevance of research, clarity of text, and compliance with the guidelines for manuscript preparation.

Manuscripts should be written preferentially in English, but texts in Portuguese or Spanish will also be considered. Studies with a broad coverage are encouraged to be submitted in English. All manuscripts should include an abstract and keywords in English and a second abstract and keywords in Portuguese or Spanish.

Authors are requested to pay attention to the instructions concerning the preparation of the manuscripts. Close adherence to the guidelines will expedite processing of the manuscript.


Manuscript Form

Manuscripts should not exceed 150 pages of double-spaced, justified text, with size 12 and source Times New Roman (except for symbols). Page format should be A4 (21 by 29.7 cm), with 3 cm of margins. The pages of the manuscript should be numbered consecutively.

The text should be arranged in the following order: Title Page, Abstracts with Keywords, Body of Text, Literature Cited, Tables, Appendices, and Figure Captions. Each of these sections should begin on a new page.

(1) Title Page: This should include the title, short title, author(s) name(s) and institutions. The title should be concise and, where appropriate, should include mention of families and/or higher taxa. Names of new taxa should not be included in titles.

(2) Abstract: All papers should have an abstract in English and another in Portuguese or Spanish. The abstract is of great importance as it may be reproduced elsewhere. It should be in a form intelligible if published alone and should summarize the main facts, ideas, and conclusions of the article. Telegraphic abstracts are strongly discouraged. Include all new taxonomic names for referencing purposes. Abbreviations should be avoided. It should not include references. Abstracts and keywords should not exceed 350 and 5 words, respectively.

(3) Body of Text: The main body of the text should include the following sections: Introduction, Material and Methods, Results, Discussion, Conclusion, Acknowledgments, and References at end. Primary headings in the text should be in capital letters, in bold and centered. Secondary headings should be in capital and lower case letters, in bold and centered. Tertiary headings should be in capital and lower case letters, in bold and indented at left. In all the cases the text should begin in the following line.

(4) Literature Cited: Citations in the text should be given as: Silva (1998) or Silva (1998:14 20) or Silva (1998: figs. 1, 2) or Silva (1998a, b) or Silva & Oliveira (1998) or (Silva, 1998) or (Rangel, 1890; Silva & Oliveira, 1998a, b; Adams, 2000) or (Silva, pers. com.) or (Silva et al., 1998), the latter when the paper has three or more authors. The reference need not be cited when authors and date are given only as authority for a taxonomic name.

(5) References: The literature cited should be arranged strictly alphabetically and given in the following format:

  • Journal Article - Author(s). Year. Article title. Jounal name, volume: initial page-final page. Names of journals must be spelled out in full.
  • Books - Author(s). Year. Book title. Publisher, Place.
  • Chapters of Books - Author(s). Year. Chapter title. In: Author(s) ou Editor(s), Book title. Publisher, Place, volume, initial page-final page.
  • Dissertations and Theses - Author(s). Year. Dissertation title. (Ph.D.Dissertation). University, Place.
  • Electronic Publications - Author(s). Year. Title. Available at: <electronic address>. Access in: date.

Tables: All tables must be numbered in the same sequence in which they appear in text. Authors are encouraged to indicate where the tables should be placed in the text. They should be comprehensible without reference to the text. Tables should be formatted with vertical (portrait), not horizontal (landscape), rules. In the text, tables should be referred as Table 1, Tables 2 and 3, Tables 2 6. Use "TABLE" in the table heading.

Illustrations: Figures should be numbered consecutively, in the same sequence that they appear in the text. Each illustration of a composite figure should be identified by capital letters and referred in the text as: Fig. 1A, Fig. 1B, for example. When possible, letters should be placed in the left lower corner of each illustration of a composite figure. Hand-written lettering on illustrations is unacceptable. Figures should be mounted in order to minimize blank areas between each illustration. Black and white or color photographs should be digitized in high resolution (300 dpi at least). Use "Fig(s)." for referring to figures in the text, but "FIGURE(S)" in the figure captions and "fig(s)." when referring to figures in another paper.

Responsability: Scientific content and opinions expressed in this publication are sole responsibility of the respective authors.

Copyrights: A concession letter of copyrights and assent should be sent to the Editor, signed by all the authors, prior to publication of the manuscript. A model is available in the home page of the Museu de Zoologia da Universidade de São Paulo.

For other details of manuscript preparation of format, consult the CBE Style Manual, available from the Council of Science Editors (

Papéis Avulsos de Zoologia and Arquivos de Zoologia are publications of the Museu de Zoologia da Universidade de São Paulo (

Always consult the Instructions to Authors printed in the last issue or in the electronic home pages: or


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