Catalogue of Dacetini and Solenopsidini ant type specimens (Hymenoptera, Formicidae, Myrmicinae) deposited in the Museu de Zoologia da Universidade de São Paulo, Brazil

Catalogue of Dacetini and Solenopsidini ant type specimens (Hymenoptera, Formicidae, Myrmicinae) deposited in the Museu de Zoologia da Universidade de São Paulo, Brazil

Mônica Antunes Ulysséa^I; Carlos R.F. Brandão^II

^IPrograma de Pós-Graduação em Sistemática, Taxonomia Animal e Biodiversidade. E-mail: [e-mail]

^IIMuseu de Zoologia, Universidade de São Paulo. Caixa Postal 42.494, 04218-970, São Paulo, SP, Brasil. E-mail: [e-mail]

ABSTRACT

The present catalogue lists ant (Formicidae) types of the Myrmicinae tribes Dacetini (Acanthognathus and Strumigenys) and Solenopsidini (Allomeurs, Carebarella, Megalomyrmex, Monomorium, Oxyepoecus, Solenopsis, Carebara and Tropidomyrmex) housed in the Formicidae collection of the Laboratório de Hymenoptera, Museu de Zoologia da Universidade de São Paulo/MZSP, Brazil. In total, the collection includes 141 types of these tribes, 41 of the Dacetini (nine holotypes and paratypes, 15 holotypes, 13 paratypes and four syntypes) and 100 of the Solenopsidini (28 holotypes and paratypes, eight holotypes, 29 paratypes, 27 syntypes, four lectotypes and paralectotypes, one lectotype and three paralectotypes), of which 37 and 89 are of still recognized species, respectively. We record label information, condition of the specimens, nomenclatural changes and type status, as well as provide indexes of the listed taxa.

Key-Words: Catalogue; Ants; Types; MZSP; Dacetini; Solenopsidini.

RESUMO

O presente catálogo lista os espécimes-tipo de espécies de formigas (Formicidae) das tribos de Myrmicinae Dacetini (Acanthognathus e Strumigenys) e Solenopsidini (Allomeurs, Carebarella, Megalomyrmex, Monomorium, Oxyepoecus, Solenopsis, Carebara e Tropidomyrmex) depositados na coleção de Formicidae do Laboratório de Hymenoptera do Museu de Zoologia da Universidade de São Paulo/MZSP, Brasil. No total, a coleção inclui 141 tipos destas tribos, sendo 41 de espécies de Dacetini (nove holótipos e parátipos, 15 holótipos, 13 parátipos e quatro síntipos) e 100 de Solenopsidini (28 holótipos e parátipos, oito holótipos, 29 parátipos, 27 síntipos, four lectótipos e paralectótipos, um lectótipo e três paralectótipos), das quais 37 e 89 ainda são ainda reconhecidas, respectivamente. São apresentados os dados de rótulos, estado de conservação dos exemplares, alterações nomenclaturais, status taxonômico e índices para os táxons catalogados.

Palavras-Chave: Catálogo; Formigas; Tipos; MZSP; Dacetini; Solenopsidini.

INTRODUCTION

The Formicidae collection housed in the Laboratório de Hymenoptera of the Museu de Zoologia da Universidade de São Paulo/MZSP is one of the most important and representative ant collections for the Neotropical region. This is the case regarding the number of types, species and specimens, as well geographic coverage (Brandão, 2000), due to the acquisition of the Walter Wolfgang Kempf and Thomaz Borgmeier collections in 1977 (Brandão, 1991). In addition, the collection had also obtained specimens belonging to the collections of Nicolas Kusnezov and William Mowry Mann. More recently, material has been added by several collaborators, including C.R.F. Brandão (the collection's curator since 1981), his students, and various other museum collaborators and associates (Esteves et al., 2011).

The ant types deposited in MZSP catalogues published thus far include fungus-growing ants (Myrmicinae: Attini) (Klingenberg & Brandão, 2005), Poneromorphs (Amblyoponinae, Ectatomminae, Heteroponerinae, Ponerinae and Proceratiinae) (Scott-Santos et al., 2008), Pseudomyrmecinae (Brandão et al., 2010) and Dorylomorphs (Aenictinae, Ecitoninae, Cerapachyinae and Leptanilloidinae) (Esteves et al., 2011).

The present catalogue deals with Dacetini and Solenopdidini ants, and is the fifth in a series aiming to list all ant types deposited at MZSP. Myrmicinae is a cosmopolite subfamily with the highest local and global species richness (Albuquerque & Brandão, 2004, 2009), including 24 tribes. Dacetini (Forel, 1892: 344) comprises eight genera and 921 recognized species (Bolton, 2012); and Solenopsidini (Forel, 1893: 164) currently includes 20 genera (two incertae sedis), 840 species and 133 subspecies (Albuquerque & Brandão, 2004, 2009; Silva et al., 2009; Bolton, 2012; Delsinne et al., 2012), representing two of the largest tribes of the subfamily.

The current situation of the Dacetini tribe deserves a comment. For the classification of ants we follow Bolton (2012), who partially accepted the propositions made by Baroni-Urbani & De Andrade (2007) for Dacetini, adopting the genus Strumigenys sensu Baroni-Urbani & De Andrade, yet still considering Basicerotini (Basiceros, Eurhopalothrix, Octostruma, Protalaridris, Rhopalothrix and Talaridris) and Phalacromyrmecini (Phalacromyrmex) as separate tribes from Dacetini (Acanthognathus, Colobostruma, Daceton, Epopostruma, Mesostruma, Microdaceton, Orectognathus and Strumigenys).

The objective of this catalogue is to provide label information and conservation status of Dacetini and Solenopsidini type specimens in the MZSP, and the current taxonomic status of the species. Our aim is to encourage further studies and also to avoid unnecessary movement of precious, sometimes unique, specimens, as proposed by the International Code of Zoological Nomenclature (1999) about institutional responsibility: "every institution in which name-bearing types are deposited should make them accessible for studies, publish lists of name-bearing types in its possession or custody, and so far as possible, communicate information concerning name-bearing types".

MATERIAL AND METHODS

For preparation of the catalogue, information from labels related to Dacetini and Solenopsidini ant types deposited at MZSP was compared to the original descriptions – which can be easily found on the Global Ant Project website (http://gap.entclub.org/taxonomists/index.html) (Bolton, 2012), the Thomaz Borgmeier catalogue and the MZSP catalogue of identifier numbers. The nomenclature of type specimens was updated following the ICZN (1999).

The combination of type names within genera and species are quoted in alphabetical order by last name, according the genus they are presently organized. We used the following abbreviations: Coll. = collection, Coll. Borgm. = T. Borgmeier Collection, Coll. Kempf = W.W. Kempf Collection, Coll. Mann = W.M. Mann Collection, Coll. Kusn. = N. Kusnezov Collection, M = worker, S = soldier (major worker), G = gyne, M = male, col(s). = collector(s) and leg. = legit. Furthermore, to indicate the conservation status of the type specimens recorded in the present catalogue, we used: A = antenna, W = wing and L = leg. We use A1 for the left antenna and A2 for the right antenna. W1 and W2 representing the anterior and posterior left wings, respectively, and W3 and W4 representing the right wings. L1, L2 and L3 represent the left legs and L4, L5 and L6 the right legs. MP is an abbreviation for Museu Paulista, of which the old Zoological section became the present MZSP in 1939.

In order to record information related to the MZSP Dacetini and Solenopsidini ant types, the following order is adopted in the present catalogue (although we do not have complete information for all entries): species name in the original combination (in alphabetical order by species within genera), author, publication year, page number, posterior taxonomic assignments, type category, number of type specimens of each sex or caste, place of collection, date, collector, original collection name, collection number or collection code and other information. All this information is taken from labels and listed as completely as they appear on the labels, compared with information in the original publication. We also comment on the conservation status of specimens. When appropriate, we indicate broken body parts glued to the paper triangle that holds the type specimen, or the ones lost.

Additional information, present in publications but not recorded on labels, is presented in brackets; for the sake of clarity, we also added further information that we were able to recover from different sources. At the end of each species entry we present, when necessary, the present valid name of the taxon corresponding to the type specimen listed. Finally, we present an index of all the valid names listed in the present catalogue and their respective synonyms or current combinations. The complete information regarding all names listed here can be found in Bolton (2012).

RESULTS

This catalogue includes information on types of 41 nominal species of Dacetini (nine holotypes and paratypes, 15 holotypes, 13 paratypes and four syntypes) and 100 of Solenopsidini (28 holotypes and paratypes, eight holotypes, 29 paratypes, 27 syntypes, four lectotypes and paralectotypes, one lectotype and three paralectotypes). Dacetini's nominal species include 156 individuals of two genera (54 of Acanthognathus and 102 of Strumigenys), of which 22 species are still valid, 14 have been already synonymized, three are junior synonyms, one was revived from the synonym and one was renamed by previous authors. There are types of 100 Solenopsidini nominal species, including 770 individuals of the eight genera (46 of Allomeurs, three of Carebarella, 146 of Megalomyrmex, eight of Monomorium, 267 of Oxyepoecus, 243 of Solenopsis, 43 of Carebara and 12 of Tropidomyrmex), of which 87 belong to still valid species, seven are considered junior synonyms, two were raised to species, one was renamed and three are considered unavailable names by previous authors.

Subfamily Myrmicinae

Tribe Dacetini

Genus Acanthognathus Mayr, 1887: 578

Acanthognathus rudis Brown & Kempf, 1969: 100. Holotype: 1W [Brazil], São Paulo, Água Funda, rotten wood, viii.1962, W.L. Brown [leg.]. Paratypes: 2W [Brazil], PR[Paraná], Bocaiúva [do Sul], v.1963, F. Plaumann [leg., Coll. Kempf] #3999. 1W [Brazil], PR[Paraná], Palmeira, v.1964, F. Plaumann [leg., Coll. Kempf] #4015. 3W [Brazil], Paraná, Volta Grande, iv.[19]49, [R.J.G.] Hertel [leg.]. 1W [Brazil, Rio de Janeiro], Itatiaya [Parque Nacional do Itatiaia], xii.[19]55, [T.] Borgmeier [leg.]. 1W [Brazil], RS[Rio Grande do Sul], Tainhas, iv.1959, F. Plaumann [leg., Coll. Kempf] #3250. 1W [Brazil], SC[Santa Catarina], Chapecó, vi.1960, F. Plaumann [leg.]. Missing L1 last tarsus. 1W [Brazil], SC[Santa Catarina], Chapecó, viii.1960, F. Plaumann [leg., Coll. Kempf] #4618. 1G [Brazil], SC[Santa Catarina], Ibicaré, vii.1959, F. Plaumann [leg., Coll. Kempf] #3126. Dealate gyne. 2W [Brazil], SC[Santa Catarina], Ibicaré, ix.1960, F. Plaumann [leg., Coll. Kempf] #3621. One worker missing L5. 2W 1G Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, x.1953, Fritz Plaumann [leg.]. Dealate gyne. 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, ii.1959, Fritz Plaumann [leg., Coll. Kempf] #3379. 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, vii.1959, Fritz Plaumann [leg.], Coll. Kempf #3100. 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, ii.1960, Fritz Plaumann [leg., Coll. Kempf] #3780. Missing A1 apical segment of the funiculus. 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, vi.1960, Fritz Plaumann [leg., Coll. Kempf] #3956. 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, vii.1961, Fritz Plaumann [leg., Coll. Kempf] #3379. 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, i.1963, Fritz Plaumann [leg., Coll. Kempf] #3744. 1G Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, vi.1963, Fritz Plaumann [leg., Coll. Kempf] #3709. Dealate gyne. 2W [Brazil], S[anta] Cat[arina], Seara, vii.1958, [F.] Plaumann [leg., Coll. Kempf] #2687. 9W [Brazil], S[ão] Paulo, Água Funda, 17.ix.1958, K. Lenko col., #608. One worker missing L2 tarsus. 1M Brazil, São Paulo, Água Funda, Jardim Botânico, wet forest, ii.[19]67, W.L. Brown [leg.]. 7W Brazil, S[ão] Paulo, Barueri, 22.xi.1958[original description says "xii"], K. Lenko leg., #708. One worker lost; one worker missing L3. 5W 1G [Brazil], SP[São Paulo], Guararema, 29.xii.1957, W.[W.] Kempf [leg., Coll. Kempf] #2120. 4W 1G [Brazil], SP[São Paulo], S[erra] da Cantareira, 1.ii.1959[original description says "i"], W.W. Kempf & V. dos Santos [leg.], Coll. Kempf #2915. Dealate gyne.

Genus Pyramica Roger, 1862a: 251

Combination in Strumigenys by Baroni-Urbani & De Andrade, 2007: 110-130.

Genus Strumigenys F. Smith, 1860: 72

Codiomyrmex reticeps Kempf, 1969: 286. Holotype: 1W Brazil, S[ão] P[aulo], Salesópolis, Est[ação] Biol[ógica de] Boracéia, 5.iii.1962, in humus, K. Lenko col. Disarticulated gaster and postpetiole glued on a paper triangle. Combination in Glamyromyrmex by Bolton, 1995: 208; in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 126.

Glamyromyrmex aztecus Kempf, 1960: 444. Paratype: 1W Mex[ico], Vera Cr[uz], Pueblo Nuevo, nr[near] Tetzonapa, rain forest, 18.viii[19]53, E.O. Wilson [leg., berlese funnel]. Missing L1. Combination in Pyramica: Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 116.

Glamyromyrmex substrictus Kempf, 1964: 68. Holotype: 1W [Brazil], PR[Paraná], Bocaiúva [do Sul], xii.1963, F. Plaumann [leg., Coll. Kempf] #3834. Disarticulated head glued on a paper triangle. Combination in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 128.

Gymnomyrmex abditus Kempf, 1960: 448. Holotype: 1W [Brazil], PR[Paraná], Rio Azul, 1000 m, x.1959, F. Plaumann [leg.], #3151. Paratypes: 4W [Brazil], PR[Paraná], Rio Azul, 1000 m, x.1959, F. Plaumann [leg.], #3151. Combination in Pyramica by Bolton, 1999: 1672, new name, P. abditivata, by Bolton 2000: 231; combination in Strumigenys by Baroni-Urbani & De Andrade, 2007: 110.

Gymnomyrmex comis Kempf, 1959: 342. Holotype: 1W, [Brazil], SC[Santa Catarina], Serra Geral, xii.1958, F. Plaumann [leg.], Coll. Kempf #3003. Missing L2 fourth and fifth tarsus, L3 femur, tibia and tarsus. Combination in Pyramica: Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 117.

Gymnomyrmex dentinasis Kempf, 1960: 450. Holotype: 1W [Brazil], PR[Paraná], Rio Azul, 1000 m, x.1959, F. Plaumann [leg.], #3150. Combination in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 118.

Gymnomyrmex minusculus Kempf, 1962: 24. Holotype: 1W Brazil, SC[Santa Catarina], Chapecó, vii.1960, [F.] Plaumann col. Combination in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 124.

Gymnomyrmex rugithorax Kempf, 1959: 339. Holotype: 1W Brazilien[Brazil, Santa Catarina], Nova Teutônia, 27º14B 52º23L, 300-500 m, xii.1958, Fritz Plaumann [leg.], Coll. Kempf #3002. Missing A2 funiculus. Combination in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 127.

Gymnomyrmex splendens Borgmeier, 1954: 279. Holotype: 1W Brazilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, x.1953, Fritz Plaumann [leg.]. Combination in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 128.

Neostruma zeteki Brown, 1959: 10. Paratype: 1W [Panama], Barro Colorado Is[land], C[anal] Z[one], iv-v.[19]42, J. Zetek [leg.], nº 4953. Missing head, L2 second to fifth tarsus. Combination in Pyramica by Bolton, 1999: 1672; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 130.

Pyramica dapsilis Bolton, 2000: 217. Holotype: 1W Brazil, SP[São Paulo], Salesópolis, E[stação] B[iológica] [de] Boracéia, 3-5.v.1996, [C.R.F.] Brandão, [D.] Agosti, [J.L.M.] Diniz, [R.] Silvestri, & [C.I.] Yamamoto [leg.]. Combination in Strumigenys by Baroni-Urbani & De Andrade, 2007: 118.

Pyramica urrhobia Bolton, 2000: 228. Paratype: 1W Trinidad, Arena Res[erve], 3.viii,[19]76, J. Noyes [leg.]. Combination in Strumigenys by Baroni-Urbani & De Andrade, 2007: 130.

Smithistruma alegrensis Brown, 1953a: 103. Holotype: 1W [Brazil], RS[Rio Grande do Sul], Porto Alegre, [Gloria], 4.x.1926, [P. Pio] Buck [S.J. leg.], Coll. Borgm. #1186. Paratypes: 5W [Brazil], RS[Rio Grande do Sul], Porto Alegre, [Gloria], 4.x.1926, [P. Pio] Buck [S.J. leg.], Coll. Borgm. #1186. One worker missing head. Junior synonym of S. emiliae Forel, 1907: 11 by Brown, 1964: 193, combination in Pyramica by Bolton 1999: 1673; combination in Strumigenys reinstated by Baroni-Urbani & De Andrade, 2007: 119.

Smithistruma cincinnata Kempf, 1974b: 419. Holotype: 1W Brazil, Amazonas, Manaus to Itacoatiara Rd[Road], km 49, 24.viii.1962, W.L. Brown [leg.]. Missing A2 and L5 second to fifth tarsus. Paratypes: 2W Brazil, Amazonas, 20-50 km N[orth] +[and] NE[Northeast] Manaus, viii-ix.1962, W.L. Brown [leg.], berlese [funnel]. One worker missing L2 second to fifth tarsus, disarticulated gaster and postpetiole glued on a paper triangle. Combination in Pyramica: Bolton, 1999: 1673; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 117.

Smithistruma sublucida Brown, 1953d: 99. Holotype: 1W, Paratype: 1W [Brazil], RS[Rio Grande do Sul], Porto Alegre, Glória, 1.viii.1926, [P. Pio] Buck [S.J. leg.], Coll. Borgm. #1196. Paratype missing L4 trochanter, femur, tibia and tarsus. Combination in Pyramica by Bolton, 1999: 1673; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 128.

Smithistruma tanymastax Brown, 1964: 183. Holotype: 1W Brazilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, [no date], Fritz Plaumann [leg.]. Only the gaster. Combination in Strumigenys by Baroni-Urbani & De Andrade, 2007: 128. Comment: In the MZSP collection there are two workers labeled as paratypes – 1W [Brazil], SC[Santa Catarina], Chapecó, 600 m, vii.1960, F. Plaumann [leg.]. Missing L4 femur, tibia and tarsus. 1W [Brazil], SP[São Paulo], S[erra da] Cantareira, 31.i.1960, [W.W.] Kempf & [V. dos] Santos [leg.], #3371 – that are not described in the original publication. We do not consider them as paratypes. Combination in Pyramica by Bolton, 1999: 1673; in Strumigenys by Baroni-Urbani & De Andrade, 2007: 128.

Strumigenys carinithorax Borgmeier, 1934: 103. Cotypes[Syntypes]: 5W 1G 3M [Surinam, ex-Dutch Guiana], Paramaribo, [Estação Experimental Agrícola, Plantação de Café, G.H.] Buenzli[Bünzli leg.]. Dealate gyne; one worker missing head; one worker missing A1/A2 funiculus.

Strumigenys clasmospongia Brown, 1953b: 2. Holotype: 1W Brazil, [Rio de Janeiro], Petrópolis, viii.22.[1938], [T.] Borgmeier [leg.], Coll. Borgm. #5884. Paratypes: 2W [Brazil, Rio de Janeiro], Petrópolis, [viii.1937], T. Borgmeier [leg.], Coll. Borgm. #5884. Junior synonym of S. louisianae Roger, 1863: 211 by Brown, 1961: 64.

Strumigenys connectens Kempf, 1958a: 59. Paratype: 1W Colombia, SA, 22.viii.[19]45, SF 19518, USNM[National Museum of Natural History] 45-16312, on Cattleya mendeli[Orchid]. Missing L1/L2/L3 trochanter, femur, tibia and tarsus, L6 tibia and tarsus, disarticulated petiole, postpetiole, gaster and head glued on a paper triangle. Combination in Pyramica by Bolton, 1999: 1673; combination in Strumigenys reinstated by Baroni-Urbani & De Andrade, 2007: 117.

Strumigenys cosmostela Kempf, 1975b: 413. Holotype: 1W Brazil, Pará, Iriboca, Pirelli [Rubber] Plantation, nr[near] Belém, BF-18, 15.viii.1962, W.L. Brown [leg.]. Missing L5 second to fifth tarsus, L6 tibia and tarsus. Paratype: 1W Brazil, Pará, nr[near] Belém, Utinga tract[State Park], BF-11, 12.viii.1962, W.L. Brown [leg.].

Strumigenys deltisquama Brown, 1957: 99. Cotype[Paratype]: 1W [Panama], Barro Colorado [Island], C[anal] Z[one], 7.i.[19]41, K.W. Cooper [leg.].

Strumigenys diabola Bolton, 2000: 533. Holotype: 1W Brazil, Amazonas, Igarapé Marianil, at R[io] Branco Rd[Road], km 5, ca. 24 km, NE[Northeast] Manaus, 22.viii.1962, W.L. Brown [leg.].

Strumigenys dubitata Bolton, 2000: 523. Paratype: 1W Costa Rica, Gste[Guanacaste] Pv[Provincia], Guanacaste N[ational] P[ark], Santa Rosa Sector, 300 m, iv.1985, J. Memmott [leg.].

Strumigenys eggersi Emery, 1890: 69. Syntype: 1W [Antilles Island, Saint Thomas Island, MZSP] #11414. Combination in Pyramica by Bolton, 1999: 1673; combinatuion in Strumigenys reinstated by Baroni-Urbani & De Andrade, 2007: 119.

Strumigenys fridericimuelleri Forel, 1886: 213. Cotype[Syntype]: 1W Brazil, [Santa Catarina], Itajahy[Blumenau, W. Müller leg., MZSP] #11.413. Combination in Pyramica by Bolton, 1999: 1673; combination in Strumigenys reinstated by Baroni-Urbani & De Andrade, 2007: 120.

Strumigenys gemella Kempf, 1974b: 416. Paratypes: 3W Colombia, Dept[Departament] Valle, Pichindé Vy[Valley], SW[Southwest] Calli, Mt. rainfor.[rain forest], rot wood, 1570 m, 22.iii.1967, R.B. Root & W.L. Brown [leg.]. Combination in Pyramica by Bolton, 1999: 1673; combination in Strumigenys reinstated by Baroni-Urbani & De Andrade, 2007: 120.

Strumigenys gytha Bolton, 2000: 553. Holotype: 1W [Brazil], SP[São Paulo], S[erra da] Cantareira, 5.iv.1973, [W.W.] Kempf [leg.], Coll. Kempf #8873.

Strumigenys infidelis Santschi, 1919: 48. Cotype[Syntype]: 1W [Argentina], La Plata, [C.] Bruch [leg., Coll. Borgm.] #3883. Species revived from the synonymy of S. louisianae Roger, 1863: 211 by Bolton, 2000: 523.

Strumigenys longispinosa Brown, 1958b: 123. Paratype: 1W [Panama], Barro Colorado [Island], C[anal] Z[one], 10.vii.[19]39, N.A. Weber [leg.], #1139, MCZ[Museum of Comparative Zoology] nº 29886. Disarticulated head glued on a paper triangle.

Strumigenys perparva Brown, 1958c: 133. Nidotypes[Paratypes]: 2W [Brazil], SP[São Paulo], Agudos, 6.iii.1955, C. Gilbert [leg., Coll. Kempf] #1376.

Strumigenys planeti Brown, 1953c: 57. Paratype: 1W Bolivia, Tumupasa, Mulf[ord] Expl[oration], iii.1922, W.M. Mann [leg.].

Strumigenys precava Brown, 1954: 196. Paratypes: 1W [Panama], Barro Colorado Is[land], C[anal] Z[one], iv-v.[19]42, [J. Zetek leg.], nº 4953, LotNo 42-15209. Missing L1/L3 tibia and tarsus, L5/L6 fifth tarsus. 3W Boliv[ia], Cachuela Esperanza, [River] Beni, iii, Mulford Bio[logical] Expl[oration] 1921-[19]22, W.M. Mann [leg.]. One worker missing L1, L2 second to fifth tarsus, L3 tarsus, L4 third to fifth tarsus.

Strumigenys princeps Kempf & Brown, 1969: 21. Holotype: 1W, Paratypes: 5W 1G Brazil, Amazonas, Ponta Negra, N[orth] Manaus, M-135[number from Brown's notebook], 1.ix.1962, W.L. Brown [& K. Lenko cols.]. One worker missing head (paratype). Paratypes: 7W 1G Brazil, Amazonas, [Ponta Negra, North] Manaus, 1.ix.1962, [W.L. Brown &] K. Lenko cols., [Coll. Kempf] #3073.

Strumigenys producta Brown, 1953b: 3. Paratype: 1W Bolivia, Beni, Rurrenabaque, x.[19]21, [Mulford Biological Exploration 1921-1922], W.M. Mann [leg.] Junior synonym of S. louisianae Roger, 1863: 211 by Bolton, 2000: 524.

Strumigenys quadrua Bolton, 2000: 557. Holotype: 1W Brazil, BA[Bahia], Ilhéus, CEPEC[Centro de Pesquisas do Cacau]-área Zoolog[setor de Zoologia], km 22 Ilhéus-Itab[una Road], x.[19]86, J. Delabie leg.

Strumigenys sanctipauli Kempf, 1958b: 556. Holotype: 1W [Brazil], SP[São Paulo], V. Estr. Santos[Old road São Paulo/Santos], Meio da Serra, 22.xii.1957, [W.W.] Kempf [leg.], Coll. Kempf #2088. Missing A1, disarticulated head glued on a paper triangle.

Strumigenys smilax Bolton, 2000: 562. Holotype: 1W Brasil, GO[Goiás], Niquelândia, Peneira Rio do Peixe, 14º17'06"S 48º55'01"W, 18-30.v.1996, [R.] Silvestre, [R.R.] Silva & [C.R.F.] Brandão col.

Strumigenys sublonga Brown, 1958a: 221. Paratypes: 3W Bolivia, Lower Rio Madidi, Mulford Biol[ogical] Expl[oration] 1921-1922, iii.1922, W.M. Mann [leg.]. One worker missing head, L1 tarsus; one worker with disarticulated L3 tibia and tarsus glued to the paper triangle.

Strumigenys thomae Kempf, 1976: 39. Holotype: 1W [Brazil], PA[Pará], Belém, 7.xi.1974, D. Dias [leg., Coll. Kempf] #13455. Paratypes: 17W [Brazil], PA[Pará], Belém, 7.xi.1974, D. Dias [leg., Coll. Kempf] #13455. One worker missing L2/L6 trochanter, femur, tibia and tarsus, L3/L4 femur, tibia and tarsus, L5 tibia and tarsus, postpetiole and gaster; one worker missing L3/L6, petiole, postpetiole and gaster; one worker missing L4 third to fifth tarsus, L5 second to fifth tarsus, L6 femur, tibia and tarsus, postpetiole and gaster; one worker missing L6; one worker missing A1; one worker missing petiole, postpetiole and gaster; one worker missing L2 femur, tibia and tarsus; one worker missing A1 funiculus, A2 last segment of the funiculus; one worker missing L1/L2 tibia and tarsus; one worker only with L1, L2 coxa and mesosoma; one worker missing head; one worker missing L6 femur, tibia and tarsus, postpetiole and gaster; one worker missing A1, part of postpetiole and gaster; one worker missing mandibles, A1/A2, L2/L3; one worker missing mandibles, clypeus, gena, A1/A2, postpetiole and gaster; one worker missing part of mandibles, A2, L4 tarsus, L5 tibia and tarsus.

Strumigenys trudifera Kempf & Brown, 1969: 18. Holotype: 1W Brazil, Pará, nr[near] Belém, Utinga tract[State Park], BF-8, 6.viii.1962, P.F. Darlington [leg.].

Tribe Solenopsidini

Solenopsis genus group

Genus Allomerus Mayr, 1878: 873

Allomerus brevipilosus Fernández, 2007a: 164. Holotype: 1W, Paratypes: 3W [Brazil], Amazonas, Rio Naué[Maués-Açu, no date, A.] Ducke [leg., in] Gleasonia nauensis. One paratype missing A2, L3 coxa, trochanter, femur, tibia and tarsus; one paratype missing A1 second to last segment of the funiculus, disarticulated L1 second to fifth tarsus glued on a paper triangle.

Allomerus decemarticulatus Mayr, 1878: 874. Syntype: 1W Brazil, AM[Amazonas], Camaná, R[io] Javari, J. Trail col. Missing A2, L1 femur, tibia and tarsus, L2/L5/L6 trochanter, femur, tibia and tarsus. Syntype designed by W.M. Wheeler, 1911: 158.

Allomerus dentatus Fernández, 2007a: 166. Holotype: 1W Venezuela, T[erritorio] F[ederal de] AM[Amazonas], Sabana El Venado, Caño Pimichín, Río Guainía, 23.xi.1953, en Tococa hirta Berg. ex-Triana, [B]. Maguire et al. [leg.], #36344, NYBG[New York Botanical Garden], WWB, #568H.

Allomerus maietae Fernández, 2007a: 167. Holotype: 1W Brasil, AM[Amazonas, Santa Isabel do] Rio Negro, betw[between] Missão Salesiana e Serra do Pirapucú [Pico da Neblina], 800-1000 m, 23.i.[19]66, in Maieta neblinensis Wurdack, N.J. Silva & U. Brazão, #60854, NYBG[New York Botanical Garden], WWB, #425H. Missing A2, L6 tarsus.

Allomerus vogeli Kempf, 1975a: 348. Holotype: 1W, Paratypes: 35W 2M [Brazil], AM[Amazonas], Mercês, 26.xii.1964, S. Vogel [leg.], #363, in Myrmidone macrosperma Martius. One worker missing L3 part of the femur, tibia and tarsus (paratype); two workers missing head (paratypes); one worker missing A1 (paratype); one male missing head, W1/W2, L1/L3.

Allomerus undecemarticulatus Fernández, 2007a: 172. Holotype: 1W Venezuela, T[erritorio] F[ederal de] AM[Amazonas], Cerro de La Neblina, Río Yatua, 1800-1900 m, 17.xi.1957, en Tococa pachystachya Wurdack, [B.] Maguire et al. [leg.], #42126, NYBG[New York Botanical Garden], WWB, #561H. Missing A1, A2 funiculus.

Genus Carebarella Emery, 1906: 137

Carebarella alvarengai Kempf, 1975a: 352. Holotype: 1G [Brazil], MT[Mato Grosso], Vila Vera, [55º30'W 12º46'S], x.1973, M. Alvarenga [leg., Coll. Kempf] #10135. Disarticulated head, M1, A2 funiculus, W1/W2 and L6 glued on a paper triangle, and W3/W4 trapped in a pin. Comment: Body parts in two pins.

Carebarella condei Borgmeier, 1937: 236. Type[Syntype]: 1M [Brazil], Espírito Santo, S[anta] Teresa, [26.vi.1928, O. Conde leg.], Coll. Borgm. #4292, [com rede]. Missing W3, A1/A2 last segment of the funiculus, disarticulated head glued on a paper triangle. Type[Syntype]: 1G [Brazil, Espírito Santo, Santa Teresa, 26.vi.1928, O. Conde leg.], Coll. Borgm. #4292, [entomological net].

Genus Epelysidris Bolton, 1987: 279

Combination in Monomorium by Fernández, 2007b: 130.

Genus Megalomyrmex Forel, 1885: 56

Leonius group

Megalomyrmex acauna Brandão, 1990: 429. Holotype: 1W [Brazil], M[ato] Grosso, Gustavo Dutra, [14º43'S 55º39'W], 25.x.[19]53, C.R. G[onçalves leg.]. Missing L6 tibia and tarsus. Paratypes: 6W [Brazil], MT[Mato Grosso], Gustavo Dutra, [14º43'S 55º39'W], 25.x.1953, C.R. Gonçalves [leg.]. One worker missing L3 second to fifth tarsus, L4 tibia and tarsus; one worker missing L2/L5/L6 tarsus; one worker missing A2 funiculus, L2 tarsus, L3 trochanter, femur, tibia and tarsus; one worker missing A2 funiculus, disarticulated L1 glued on a paper triangle; one worker missing L2, disarticulated head and frist segment of the thorax glued on a paper triangle. 1W [Brazil], Mato Grosso, Chapada, [15º26'S 55º45'W], vii.1960, C. Amann col. Missing A2 funiculus.

Megalomyrmex bidentatus Fernández & Baena, 1997: 112. Paratype: 1W Colombia, Nariño, Barbacoas, Tajadas, 01º17'N 78º15'W, 1000 m, ago[sto.19]94, F. Escobar [leg.], #446. Missing L2/L6 last tarsus, L5 trochanter, femur, tibia and tarsus.

Megalomyrmex cyendyra Brandão, 1990: 433. Paratypes: 1W Colombia, Valle [del Cauca], Diguá[Dagua, Parque Nacional Natural Farollones de Cali, El] Queremal, Rio San Juan, [03º31'N 76º43'W], 1300 m, 28.vii.[19]73, W.L. Brown [leg.]. Missing A1. 2W Colombia, Valle del Cauca, Park [Nacional Natural] Farollones de Cali, El Topazio, [Rio] Pance, [03º20'N 76º43'W], 31.xii.1981, J.E. Lattke col.

Megalomyrmex glaesarius Kempf, 1970: 355. Holotype: 1W Peru, [Cajamarca], Llama, [06º31'S 79º08'W], 2350 m, [no date, W.] Weyrauch [leg.], #1069. Missing L6 femur, tibia and tarsus. Paratypes: 26W Peru, [Cajamarca], Llama, [06º31'S 79º08'W], 2350 m, [no date, W.] Weyrauch [leg.], #1069. One worker missing L5 coxa, trochanter, femur, tibia and tarsus; two workers with disarticulated postpetiole and gaster glued on a paper triangle; one worker missing A1, L1 tibia and tarsus; one worker missing L6 femur, tibia and tarsus; one worker missing A2, L5 femur, tibia and tarsus.

Megalomyrmex leoninus Forel, 1885: 57. Type[Paralectotype]: 1W Colombia, Coll. Forel. Missing A1/A2, disarticulated L2 glued on a paper triangle, and L3 glued in the gaster.

Megalomyrmex pacova Brandão, 1990: 437. Paratype: 1W Ecuador, 26.iv.1957, in bananas [specimen came from banana shipments arrived at U.S. Quarentine ports – Charleston, South Carolina; Brownsville, Texas; and New York], F. Perlmutter [leg.].

Megalomyrmex timbira Brandão, 1990: 439. Holotype: 1W, Paratype: 1W Venezuela, Táchira, Palmira, La Flautera, 1200 m, 8.viii.1983, J. Lattke col. Paratype with disarticulated L4, postpetiole and gaster glued on a paper triangle.

Modestus group

Megalomyrmex ayri Brandão, 1990: 420. Holotype: 1W Brasil, MT[Mato Grosso], Utiariti, Rio Papagaio, 325 m, vii.1961, K. Lenko col., #1731. Paratypes: 3W 1G Brasil, MT[Mato Grosso], Utiariti, Rio Papagaio, 325 m, vii.1961, K. Lenko col., #1731. One worker missing A1, L2/L3 trochanter, femur, tibia and tarsus, disarticulated head, mandibles and mouth apparatus glued on a paper triangle; one worker with disarticulated head, mandibles, mouth apparatus and A1/A2 glued on a paper triangle; one worker with disarticulated head, mandibles and mouth apparatus glued on a paper triangle; dealate gyne. 3W [Brazil], MT[Mato Grosso], Sinop, 55º37'W 12º31'S, x.1974, [M.] Alv[arenga] & [O.] Roppa [leg.], #12516. One worker missing L2 tibia and tarsus; one worker missing L6 second to fifth tarsus. 3W 1G Brasil, Mato Grosso, Diamantino, Faz[enda] Junqueira Vilela, [14º25'S 56º26'W], 3 km E[ast] casa, for., 17-18.vii.1973, W.L. Brown [leg.]. One worker missing L5 tibia and tarsus; one worker missing L1 femur, tibia and tarsus; dealate gyne. 4W 2G Brasil, Rondônia, Vilhena, Est[rada] V[elha] Colorado, [Chácara Pedro], mata [perto de igarapé], litter, ninho, 31.vii.1988, Ana Yoshi Harada [leg.]. Dealate gynes; mandibles from other specimen glued on a paper triangle.

Megalomyrmex caete Brandão, 1990: 422. Paratypes: 1W 1G Ecuador, Prov[incia] Guayas, 3 km SW[Southwest] Bucay, mt-base for., 24.vii.[19]73, W.L. Brown [leg.]. One worker missing M2, disarticulated mouth apparatus glued on a paper triangle; dealate gyne missing A2 funiculus.

Megalomyrmex cupecuara Brandão, 1990: 423. Paratype: 1M Colombia, [Departament] Valle [del Cauca], 4 km E[ast] Sabaletos, Trop[ical] rain forest, 18.x.1975, R.C. Wilkerson col., malaise trap, Venezuela-Inst[ituto de] Zool[ogía] Agricola, Fac[uldad de] Agronomia, U[niversidad] C[entral de] V[enezuela, Campus] Maracay. Missing part of A2. Paratypes: 1W 1M Colombia, [Departament] Valle [del Cauca], Munic[ipio] Buenaventura, Anchicayá, rainfor[est], 200 m, 17-19.vi.1971, W.L. Brown [leg.]. Male missing part of A1/A2. 1M Ecuador, [Provincia Pichincha], 16 km SE[Southeast] S[anto] Domingo de los Colorados, Tinalandia, vi.1975, S. [Peck] & J. Peck. [leg.].

Megalomyrmex goeldii Forel, 1912: 14. Paralectotype: 1W Brasil, RJ[Rio de Janeiro], Colônia Alpina[Teresópolis, no date, E.A.] Goeldi col.

Megalomyrmex iheringi Forel, 1911: 304. Cotype[Lectotype]: 1W [Brazil, São Paulo], A[lto] da Serra[Paranapiacaba, Santo André, 23º47'S 46º19'W, 22.viii.1910, H.F.A.] v. Jheringi [leg., Coll. Borgm.] #2093. Cotypes[Paralectotypes]: 3W [Brazil, São Paulo], A[lto] da Serra[Paranapiacaba, Santo André, 23º47'S 46º19'W, 22.viii.1910], H.[F.A.] v. Jheringi [leg., Coll. Borgm.] #2093. 7W [Brazil], SP[São Paulo], Alto da Serra[Paranapiacaba, Santo André, 23º47'S 46º19'W, 22.viii.1910], H.[F.A.] v. Jheringi [leg., MZSP] #15527. 4W [Brazil, São Paulo, Santo André, Paranapiacaba], Parque Cajuru[Reserva Biológica do Alto da Serra de Paranapiacaba, 27.ix.1909, H.] Lüderwaldt [leg., MZSP] #14096. Cotypes[Not types]: 1G [Brazil, São Paulo], A[lto] da Serra[Paranapiacaba, Santo André, 22.viii.1910, H.F.A.] v. Jheringi [leg., Coll. Borgm.] #2093. Dealate gyne. 1G [Brazil], SP[São Paulo], Alto da Serra[Paranapiacaba, Santo André], 22.viii.1910, [H.F.A.] v. Jheringi [leg.], #2095. Dealate gyne. Comment: These two gynes labelled as cotype were not described by Forel, who studied only workers, thus we do not consider them as types.

Megalomyrmex weyrauchi Kempf, 1970: 357. Holotype: 1W Peru, [Cajamarca], Llama, [06º31'S 79º08'W], 2350 m, [no date, W.] Weyrauch [leg., Weyrauch accession number] #1070. Paratypes: 20W Peru, [Cajamarca], Llama, [06º31'S 79º08'W], 2350 m, [no date, W.] Weyrauch [leg., Weyrauch accession number] #1070. Two workers missing A1; one worker missing head; one worker missing A2, L6 tarsus, disarticulated mandibles and mouth apparatus glued on a paper triangle.

Pusillus group

Megalomyrmex drifti Kempf, 1961: 504. Holotype: 1W, Paratypes: 4W Surinam, Dirkshoop, [40 km west of Paramaribo – capital of Surinam], DIIIcd[soil sample taken in primary forest], x.1959, I[J.] v[an] d[er] Drift col. One worker missing petiole, postpetiole and gaster (paratype).

Megalomyrmex gnomus Kempf, 1970: 359 Holotype: 1W [Brazil], MT[MS/Mato Grosso do Sul], Corumbá, 18.x.[19]53, C.R. G[onçalves leg.]. Paratypes: 2W [Brazil], MT[MS/Mato Grosso do Sul], Cuyabá[Corumbá], 18.x.[19]53, C.R. G[onçalves leg.]. One worker missing A1, disarticulated petiole, postpetiole and gaster glued on a paper triangle. Comment: In the original description all types were designed from the same locality, Corumbá, and Dr. Gonçalves confirmed this information (Brandão, 1990).

Megalomyrmex incisus M.R. Smith, 1947: 102. Paratypes: 3W [Panama], Barro Colorado Is[land], C[anal] Z[one], vi-x.1943, [J.] Zetek [leg.], nº 5105, Lot. 43-16534, USNM[National Museum of Natural History] 58040. One worker missing A2; one worker with disarticulated head and missing A1; one worker missing L1/L2 last tarsus, disarticulated mandibles, mouth apparatus and A1 glued on a paper triangle.

Megalomyrmex miri Brandão, 1990: 452 Paratype: 1W Costa Rica, Zent, iii.1924, W.M. Mann [leg.], Coll. Mann 1954. Missing A1 and L6 last tarsus.

Silvestrii group

Megalomyrmex cuatiara Brandão, 1990: 440. Holotype: 1W Colombia, Amazonas, 7 km NW[Northwest] Leticia, [04º09'S 69º57'W], forest, litter, 20-25.ii.1972[original publication says "iii"], S. [Peck] & J. Peck [leg.], #250, berl[ese funnel]. Missing L2 tibia and tarsus. Paratypes: 1G Colombia, Amazonas, 7 km NW[Northwest] Leticia, [04º09'S 69º57'W], forest, litter, 20-25.ii.1972[original publication says "iii"], S. [Peck] & J. Peck [leg.], #250, berl[ese funnel]. Dealate gyne. 2W 1G Colombia, Putumayo, Villa Garzón, [01º02'N 76º37'W], 14.viii.[19]77, B. Jackson [leg.]. One worker missing L2 last tarsus. 1W Colombia, Meta, Villavicencio, [04º10'N 73º37'W], forest, litter, 1-4.iii.1972, S. [Peck] & J. Peck [leg.], #233, berl[ese funnel]. Missing postpetiole and gaster, disarticulated mandibles and A1 glued on a paper. 1G Colombia, Meta, Quebrada Susamuko, 23 km NW[Northwest] Villavivencio, for[est], litter, 1000 m, 5.iii. 1972, S. [Peck] & J. Peck [leg.], #B-234.

Megalomyrmex mondabora Brandão, 1990: 442. Paratypes: 1W Costa Rica, 3-5 km E[ast] Turrialba, R[io] Reventazón, [09º45'N 83º41'W], 18-22.i.1973, W.L. Brown [leg.]. Missing L4 last tarsus, disarticulated A1, mandibles and mouth apparatus glued on a paper. 1W [Brazil], RO[Rondônia], Vilhena, [12º43'S 60º07'W], xi.1973, M. Alvarenga [leg.], #10221. Missing A1, L1/L2/L3 femur, tibia and tarsus, L4 last tarsus.

Megalomyrmex mondaboroides Longino, 2010: 49. Paratype: 1W Panama, [Provincia] Panama, El Lano, 9.279555ºN 78.9615ºW, forest, in underground nest of Apterostigma goniodes Lattke, 1997, 30.iii.2011, H. Fernandez-Marin [leg.], #HF010330-50. Comment: One worker of host species, A. gonoides, in the same pin.

Megalomyrmex nocarina Longino, 2010: 51. Paratype: 1W Costa Rica, [Heredia, Cantarrana], 11 km ESE[East-Southeast] La Virgen, 10.34528[ºN] 84.05778[ºW], wet forest, leaf litter, 300-500 m, 26.ii.2007, [no leg., Proyecto] TEAM AMI-4-W-123-09.

Megalomyrmex piriana Brandão, 1990: 443. Paratype: 1W Ecuador, Prov[incia] Pichincha, Tinalandia, 16 km SE[Southwest] S[anto] Domingos de los Colorados, [00º18'S 79º04'W], 4.vi.1976, S. [Peck] & J. Peck [leg.]. Missing L1, L2 tibia and tarsus.

Megalomyrmex sjöstedti Wheeler, 1925: 30. Types[Syntypes]: 3W Peru, Llinquipata, [13º49'S 70º38'W], N. Holmgrem [leg.]. Junior synonym of Megalomyrmex silvestrii Wheeler, 1909: 235 by Kempf & Brown, 1968: 97.

Megalomyrmex symmetochus Wheeler, 1925: 168. Lectotypus[Paralectotypes]: 1W 1G [Panama], Barro Colorado I[sland], C[anal] Z[one, 09º10'S 79º51'W], 30.vii.1924, W.M. Wheeler [leg.], nº 790. Dealate gyne, two wings glued on a paper. 1W [Panama], Barro Colorado I[sland], C[anal] Z[one, 09º10'S 79º51'W], 24.i.1924, W.M. Wheeler [leg.], nº 769. Disarticulated A1, mandibles and mouth apparatus glued on a paper. 1W [Panama], Mt[Mount] Hope, nr[near] Colón, C[anal] Z[one], 24.vii.1924, W.M. Wheeler [leg.], nº 616.

Megalomyrmex tasyba Brandão, 1990: 446. Holotype: 1W Ecuador, Prov[incia] Napo, Limoncocha, 00º24'S 76º36'W, 3.viii.1972, C.W. Rettenmeyer [leg.], #4911. Missing A1.

Megalomyrmex wettereri Brandão, 2003: 155. Paratypes: 2W 1G Panama, Prov[incia] Panama, km 7 of El Llano Carti Suitupo Road, 09º18'02"N 78º57'31"W, 7.vii.1998, U. Mueller [leg.], #980607-01. Dealate gyne missing A1 funiculus. 8W Costa Rica, [Heredia], La Selva, [10º26'N 84º01'W], em ninho Trachymyrmex bugnioni, 2.iii.1993, J. Wetterer col. One worker missing A1, L2 trochanter, femur, tibia and tarsus; one worker missing A1.

Genus Monomorium Mayr, 1855: 452

Megalomyrmex bicolor Ettershank, 1965: 55. Paratype: 1W Chile, [Provincia Coquimbo], nr[near] La Serena, Pachon Peak[Cerro Pachón], 8500ft[feet, only 25 miles from the crest of the Andes], 18.ii.1964, E. Ackerman [leg.], MCZ[Museum of Comparative Zoology]. Combination in Nothidris by Snelling, 1975: 3, in Monomorium by Heterick, 2006: 79 and by Fernández, 2007b: 131. Junior secondary homonym of Monomorium bicolor Emery, 1877: 368. New name proposed for M. bicolor (Ettershank, 1965), as M. chilensis, by Fernández, 2007b: 132.

Monomorium (Notomyrmex) bidentatum piceonigrum Borgmeier, 1948: 468. Type[Syntype]: 1W Arg[entina, Neuquén], Hua Hum[extremo oeste del Lago Lacar, cerca de La frontera chilena], xii.[19]47, Hayward-Willink & [N.] Kusnezov [leg.], #1775. Combination in Antichthonidris by Snelling, 1975: 6, junior synonym of A. bidentata by Bolton, 1995: 67, combination revised and junior synonym of Monomorium bidentatum Mayr, 1887: 616 by Fernández, 2007b: 131.

Monomorium (Notomyrmex) denticulatum inerme Borgmeier, 1948: 469. Type[Syntype]: 1W Arg[entina, Neuquén], Hua Hum[extremo oeste del Lago Lacar, cerca de La frontera chilena], xii.[19]47, Hayward-Willink & [N.] Kusnezov [leg.], #1775. Combination in Antichthonidris by Snelling, 1975: 6, junior synonym of A. denticulata by Bolton, 1995: 67, combination revised and junior synonym of Monomorium denticulatum Mayr, 1887: 614 by Fernández, 2007b: 131.

Monomorium inusuale Fernández, 2007b: 135. Paratypes: 2W Ecuador, Prov[incia] Napo, Limoncocha, 00º24'S 76º36'W, 280 m, 13.viii.1973, M. Rettenmeyer [leg.], #68.

Monomorium minimum cyaenum Wheeler, 1914: 43. Paratypes: 3W Mexico, Hidalgo, Guerrero Mill, [under stones in rather damp places on the sides of canyons, emersoni spring or summer, 1913, W.M.] Mann [leg.], Coll. Mann #1954. One worker missing postpetiole and gaster. Raised to species by DuBois, 1981: 35 and senior synonym of M. emersoni Gregg, 1945: 66 by DuBois, 1986: 82.

Genus Nothidris Ettershank, 1966: 105

Combination in Monomorium by Fernández, 2007b: 130.

Genus Oxyepoecus Santschi, 1926: 6

Rastratus group

Martia daguerrei Santschi, 1933: 111. Type[Lectotype]: 1W [Argentina], Provincia Buenos Aires, Rosas F.C. Sud, [35º57'S, 58º56'W, no date], Juan B. Daguerre [leg.], #1904-666. Missing A2. Combination in Oxyepoecus by Ettershank, 1966: 146 and designation of lectotype by Kempf, 1974a: 486.

Monomorium rastratum var. luederwaldti Forel, 1913: 219. Syntypes: 12W 1G [Brazil], SP[São Paulo], Salto Grande[Paranapanema River (Lüderwaldt, 1918)], ii.1911, H. Lüderwaldt [leg.], MP[MZSP] #15997. One worker missing A2 funiculus; one worker missing L3 trochanter, femur, tibia and tarsus; dealate gyne. Combination in Oxyepoecus by Ettershank, 1966: 146; junior synonym of O. rastratus (Mayr, 1887: 615) by Kempf, 1974a: 498. Comment: The MZSP has a specimen labeled with the number #15997, but without information on its type status. As the original description does not mention the precise number of types, we do not consider this last specimen a type.

Oxyepoecus myops Albuquerque & Brandão, 2009: 295. Holotype: 1W Brazil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia, 23º39'S, 45º53'W, leaf litter], 5-7.ix.1997[original description says "vii"], C.I. Yamamoto col., winkler. Paratypes: 5W 1G Brazil, SP[São Paulo], Salesópolis, EB[Estação Biológica de] Boracéia, 23º31'56"S 45º50'47"[W], 3-5.v.1996, [C.R.F.] Brandão, [D.] Agosti, [J.L.M.] Diniz, [R.] Silvestri, & [C.I.] Yamamoto [leg.], winkler. One worker with disarticulated head glued on a paper triangle and missing A2; dealate gyne. 16W Brazil, SP[São Paulo], Salesópolis, EB[Estação Biológica de] Boracéia, 2-6.v.1997, [D.] Agosti, [C.R.F.] Brandão & [C.I.] Yamamoto col., winkler. One worker missing L1 trochanter, femur, tibia and tarsus; one worker missing A2, A1 last segment of the funiculus; one worker missing postpetiole and gaster; one worker missing L3 tibia and tarsus; one worker missing head; one worker missing L1 trochanter, femur, tibia and tarsus, L2/L3 tibia and tarsus; one worker just with L5/L6, petiole, postpetiole and gaster; one worker missing A1 funiculus, L1 trochanter, femur, tibia and tarsus. 2W Brazil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia, 23º39'S, 45º53'W, leaf litter], 5-7.ix.1997[original description says "vii"], C.I. Yamamoto col., winkler. One worker missing L5 tarsus. 11W 2G Brazil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia], 12-17.vi.1997, B.H. Dietz & C.I. Yamamoto col., winkler. One worker missing A1; two workers missing A1 funiculus; one worker just with L5/L6; dealate gynes.

Oxyepoecus plaumanni Kempf, 1974a: 492. Holotype: 1W [Brazil], RS[Rio Grande do Sul], Erechim, vii.1960, F. Plaumann [leg., Coll. Kempf] #8097. Paratypes: 12W [Brazil], RS[Rio Grande do Sul], Erechim, vii.1960, F. Plaumann [leg., Coll. Kempf] #8097. One worker lost; one worker just with A2/L4. 2W [Brazil], RS[Rio Grande do Sul], Erechim, vii.1960, F. Plaumann [leg., Coll. Kempf] #9930. One worker lost; one worker missing A2/L1/L2/L3, L6 tarsus, disarticulated head glued on a paper triangle and coated with gold for SEM examination. 1W [Brazil], RS[Rio Grande do Sul], Tainhas, iv.1959, F. Plaumann [leg., Coll. Kempf] #3247. Worker lost. 1W [Brazil], RS[Rio Grande do Sul], B[arão de] Cotejipe, vii.1960, F. Plaumann [leg., Coll. Kempf] #3765. 1W [Brazil], RS[Rio Grande do Sul], Boqueirão, 600 m, ix.1960, F. Plaumann [leg., Coll. Kempf] #3611. Missing L3 trochanter, femur, tibia and tarsus. 1W [Brazil], PR[Paraná], Porto Vitória, 26º10'[S] 51º12'[W], 700 m, x.1959, F. Plaumann [leg., Coll. Kempf] #3209. 1W [Brazil], PR[Paraná], Rio Azul, 1000 m, x.1959, F. Plaumann [leg., Coll. Kempf] #3189. 2W [Brazil], SC[Santa Catarina], Chapecó, vi.1960, F. Plaumann [leg.]. 1W [Brazil], SC[Santa Catarina], Chapecó, viii.1960, F. Plaumann [leg., Coll. Kempf] #8331. 2W [Brazil], SC[Santa Catarina], Chapecó, viii.1960, F. Plaumann [leg., Coll. Kempf] #9883. One worker lost. 1W [Brazil], SC[Santa Catarina], Chapecó, vii.1960, F. Plaumann [leg., Coll. Kempf] #9911. 2W [Brazil], SC[Santa Catarina], Ibicaré, ix.1960, F. Plaumann [leg., Coll. Kempf] #8781. 1G [Brazil], SC[Santa Catarina], Passo Bormann, xii.1957, [F.] Plaumann [leg.]. Dealate gyne. 2W [Brazil], SC[Santa Catarina], Nova Teutônia, vi.1960, F. Plaumann [leg., Coll. Kempf] #8091. One worker lost. 1G [Brazil], SC[Santa Catarina], Nova Teutônia, xii.1972, F. Plaumann [leg., Coll. Kempf] #8557. Missing gaster. 1G [Brazil], SC[Santa Catarina], Nova Teutônia, xii.1972, F. Plaumann [leg., Coll. Kempf] #8383. Dealate gyne. Comment: In the original description, Kempf (1974a) did not consider as paratypes three workers from [Brazil], SC[Santa Catarina], Linha Facão, v.1957, F. Plaumann [leg.], however these specimens, all in the same pin, are deposited in the MZSP and labeled as paratypes, but we do not consider them as types. We were not able to locate coordinates for Linha Facão, SC.

Oxyepoecus regularis Ulysséa & Brandão, 2012: 169. Holotype: 1W Brasil, Bahia, Milagres, Caatinga arbórea, serapilheira, winkler, área 1, S12º54.542' W39º51.279', 23.x.2010, M.A. Ulysséa, A.M. Medina & E.M. Campos leg. Paratypes: 19W Brasil, Bahia, Milagres, Caatinga arbórea, serapilheira, winkler, área 3, S12º54.411' W39º50.863', 17.i.2011, M.A. Ulysséa, A.M. Medina & E.M. Campos leg. One worker missing L6 tibia and tarsus; one worker missing L6 trochanter, femur, tibia and tarsus. 7W Brasil, Bahia, Milagres, Caatinga arbórea, serapilheira, winkler, área 3, S12º54.411' W39º50.863', 9.xi.2010, M.A. Ulysséa, A.M. Medina & E.M. Campos leg. One worker missing L4 trochanter, femur, tibia and tarsus; one worker missing L1 tibia and tarsus; one worker missing A2, L1 tibia and tarsus. 12W Brasil, Bahia, Milagres, Caatinga arbórea, serapilheira, winkler, área 1, 12º54.542'S 39º51.279'W, 10.i.2011, M.A. Ulysséa, A.M. Medina & E.M. Campos leg. One worker missing A1, L1 tibia and tarsus; one worker missing L1 trochanter, femur, tibia and tarsus; one worker missing A1, L4 trochanter, femur, tibia and tarsus; one worker with disarticulated head glued on a paper triangle. 5W 1G Brasil, Bahia, Milagres, Caatinga arbórea, serapilheira, winkler, área 2, S12º54.294' W39º52.083', 10.i.2011, M.A. Ulysséa, A.M. Medina & E.M. Campos leg. One worker missing A1. 1W Brasil, Bahia, Boa Vista do Tupim, Mata Seca, S12º39'36" W40º36'32", 25.i.2011, L.S.S.R. Macêdo, J.J. Resende, C.B.S. Galheigo & E. da C. Menezes leg.

Oxyepoecus reticulatus Kempf, 1974a: 502. Holotype: 1W [Brazil], SC[Santa Catarina], N[ova] Teutônia, viii.1952, F. Plaumann [leg., Coll. Kempf] #5954. Missing head, L1/L3. Paratypes: 11W [Brazil], SC[Santa Catarina], N[ova] Teutônia, viii.1952, F. Plaumann [leg., Coll. Kempf] #5954. One worker coated with gold for SEM examination missing A2, L1/L3; one worker missing petiole, postpetiole and gaster. 1W [Brazil], PR[Paraná], Buaragi[Guaragi], v.1964, F. Plaumann [leg., Coll. Kempf] #4008. 1W [Brazil], PR[Paraná], Guaragi, 25º16'[S] 50º14'[W], 1000 m, v.1964, F. Plaumann [leg., Coll. Kempf] #4580. 1G [Brazil], PR[Paraná], Mariópolis, [no date], F. Plaumann [leg.]. Dealate gyne with disarticulated postpetiole and gaster glued on a paper triangle. 1W [Brazil], PR[Paraná], Rio Azul, 1000 m, x.1959, F. Plaumann [leg., Coll. Kempf] #3188. 2W [Brazil], PR[Paraná], Rolândia, 6.iv.1955, [W.]W. Kempf [leg., Coll. Kempf] #1414. 6W [Brazil], S[anta] Cat[arina], Chapecó, v.1957, F. Plaumann [leg.]. 4W [Brazil], SC[Santa Catarina], Chapecó, vi.1960, F. Plaumann [leg.]. One worker coated with gold for SEM examination missing postpetiole and gaster. 3W [Brazil], SC[Santa Catarina], Chapecó, vii.1960, F. Plaumann [leg., Coll. Kempf] #9910. 1W [Brazil], SC[Santa Catarina], Concórdia, vii.1958, [F.] Plaumann [leg.]. Missing head. 6W [Brazil], SC[Santa Catarina], Linha Facão, v.1957, F. Plaumann [leg.]. 6W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, viii.1952, Fritz Plaumann [leg.]. 1W [Brazil], SC[Santa Catarina], Passo Bormann, xii.1957, F. Plaumann [leg.]. One worker missing A1. 8W [Brazil], S[ão] P[aulo], Agudos, 25.iii.1955, [W.]W. Kempf [leg., Coll. Kempf] #1405. 2W [Brazil], S[ão] P[aulo], Agudos, 4.i.1956, [W.]W. Kempf [leg., Coll. Kempf] #1552. One worker lost. 2W [Brazil], S[ão] P[aulo], Agudos, 8.i.1956, [W.]W. Kempf [leg., Coll. Kempf] #1560. 1W [Brazil], SP[São Paulo], Agudos, xi.1959, C. Gilbert [leg., Coll. Kempf] #3218. Worker lost. 1W [Brazil], SP[São Paulo], Campos de Jordão, 16.x.1956, [W.]W. Kempf [leg., Coll. Kempf] #1601. 6W [Brazil, São Paulo], V. Estr. Santos[Old road São Paulo/Santos], Meio da Serra, 8.viii.1960, [W.W.] Kempf [leg.]. Comment: Original description reports that all specimens were sampled with berlese funnels.

Oxyepoecus rosai Albuquerque & Brandão, 2009: 304. Holotype: 1W Brasil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia], 20-26.x.1997, C. Klingenberg & C.I. Yamamoto col., sardinha[attracted to sardine baits]. Paratypes: 16W 1G Brasil, SP[São Paulo], Salesópolis, EB[Estação Biológica de] Boracéia, 2-6.v.1997, D. Agosti, C.R.F. Brandão & C.I. Yamamoto col., winkler. One worker missing L1 trochanter, femur, tibia and tarsus; one worker missing A1; two workers missing L2 trochanter, femur, tibia and tarsus; two workers missing head; dealate gyne. 2W Brazil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia], 5-7.vii.1997, C.I. Yamamoto col., winkler. One worker missing head. 2W 1G Brasil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia], 5-7.ix.1997, C.I. Yamamoto col., winkler. Dealate gyne with disarticulated A1 glued on paper triangle and missing A1 last segment of the funiculus. 15W Brasil, SP[São Paulo], Salesópolis, EBB[Estação Biológica de Boracéia], 12-17.vi.1997, B.H. Dietz & C.I. Yamamoto col., winkler. One worker missing A1, L3 trochanter, femur, tibia and tarsus; one worker missing A1/A2 funiculus; one worker missing L1 trochanter, femur, tibia and tarsus; one worker missing A1 funiculus; one worker missing L2 trochanter, femur, tibia and tarsus; one worker missing L3 tarsus; one worker coated with gold for SEM examination missing L2 femur, tibia and tarsus.

Vezenyii group

Martia inquilina Kusnezov, 1952a: 720. Paratypes: 3W Argentina, Tucumán, Jardín de la Fundación Miguel Lillo, en nido de Pheidole radoszkowskii Mayr, 1884: 35, 6.i.1948, N. Kusnezov [leg.], #111. 2W Argentina, Tucumán, Quebrada Cainzo, [near Tafí Viejo], en nido de Pheidole silvestrii Emery, 1906: 146, 8.iv.1948, N. Kusnezov [leg.], #1590. Diagnosis in key (Kusnezov, 1952a), combination in Oxyepoecus by Ettershank, 1966: 146 and senior synonymy of O. turgidus Kempf, 1969: 278 by Kempf, 1974a: 495.

Monomorium punctifrons Borgmeier, 1928: 39. Lectotype: 1W [Brazil], PR[Paraná], Rio Negro, iv.1925, [P.]M. Witte [leg.], Coll. Borgm. #424. Paratypes[Paralectotypes]: 15W [Brazil], PR[Paraná], Rio Negro, iv.1925, [P.]M. Witte [leg.], Coll. Borgm. #424. One worker missing A2, L6 tibia and tarsus; one worker with disarticulated postpetiole and gaster. Combination in Oxyepoecus by Ettershank, 1966: 146 and designation of lectotype and paralectotype by Kempf, 1974a: 495.

Oxyepoecus Browni Albuquerque & Brandão, 2004: 60. Holotype: 1W Brasil, MG[Minas Gerais], Atalaia, [18º03'S 41º06'W], 27.i.[19]94, I[van] Cardoso [leg.], #4772. Specimen coated with gold for SEM examination.

Oxyepoecus bruchi Santschi, 1926: 6. Lectotype: 1G, Paratypes[Paralectotypes]: 2G [Argentina], Sierras de Córdoba[Provincia Córdoba], La Granja, Alta Gracia, [31º39'S 64º25'W, no date], C. Bruch leg, #1694. Dealate gynes; one gyne missing head, postpetiole and gaster (paralectotype). Comment: One worker of host species, Pheidole obtusopilosa Mayr, 1887: 586, mounted in the same pin as the lectotype. Senior synonym of O. minuta (Kusnezov, 1952a: 721) and lectotype designation by Kempf, 1974a: 478.

Oxyepoecus crassinodus Kempf, 1974a: 482. Holotype: 1W [Brazil], PR[Paraná], Bocaiúva [do Sul, 25º12'S 49º07'W], xii.1963, F. Plaumann [leg., Coll. Kempf] #8108. Missing L3 tibia and tarsus. Paratypes: 3W [Brazil], PR[Paraná], Bocaiúva [do Sul, 25º12'S 49º07'W], xii.1963, F. Plaumann [leg., Coll. Kempf] #8108. 1W [Brazil], SC[Santa Catarina], N[ova] Teutônia, [27º02'S 52º24'W], xii. 1958, F. Plaumann [leg., Coll. Kempf] #8308.

Oxyepoecus ephippiatus Albuquerque & Brandão, 2004: 64. Paratypes: 1W 1G Brasil, AM[Amazonas], Manaus, 29.ix.[19]93, A.B. Casimiro [leg.], #2306. Comment: Worker incomplete (just mesosoma, petiole and L4 present) coated with gold for SEM examination.

Oxyepoecus kempfi Albuquerque & Brandão, 2004: 67. Holotype: 1W Brasil, PI[Piauí], 10 km N[orth] Corrente, Faz[enda] Maracujá, [10º26'S 45º09'W], folhiço, 23-27.xi.1991, C.R.F. Brandão [leg.]. Missing L2 part of femur, tibia and tarsus, disarticulated postpetiole and gaster glued on a paper triangle. Paratype: 1W Brasil, PI[Piauí], 10 km N[orth] Corrente, Faz[enda] Maracujá, [10º26'S 45º09'W], folhiço, 23-27.xi.1991, C.R.F. Brandão [leg.]. Missing A1, L1/L2/L3/L4/L5 tibia and tarsus, L6, disarticulated postpetiole and gaster glued on a paper triangle.

Oxyepoecus longicephalus Albuquerque & Brandão, 2004: 69. Holotype: 1W Brasil, RS[Rio Grande do Sul, Itatí], CPCN[Centro de Pesquisa e Conservação da Natureza] Pró-Mata, 29º30'S 50º10'W, Mata Atlântica, 670-690 m, litter, 28.v.1997, J. Ketterl [leg.], winkler. Paratypes: 1W Brasil, RS[Rio Grande do Sul, Itatí], CPCN[Centro de Pesquisa e Conservação da Natureza] Pró-Mata, 29º30'S 50º10'W, Araucaria Forest, 900 m, litter, 15.iv.[19]97, J. Ketterl [leg.], winkler. Missing A1/A2, L2 femur, tibia and tarsus, L3/L6 tibia and tarsus. 1W Brasil, SP[São Paulo], Cunha, P[arque] E[stadual] Serra do Mar, 23º15'03"S 45º00'26"W, 21-22.iv.2001, A.A. Tavares & R.R. Silva cols., winkler.

Oxyepoecus quadratus Albuquerque & Brandão, 2004: 73. Holotype: 1W Ecuador, Cuyabeno, [00º16'S 75º53'W], 12.x-5.xi.[19]94 [original description says "27.i.1994"], J.P. Caldwell [leg.], #10523. Missing A1/A2 funiculus, L5/L6 tarsus, coated with gold for SEM examination. Paratype: 1W Peru, Huánuco, Panguana Biol[ogical] Stat[ation], Rio Yuyapidis, 09º37'S 74º56'W, innundation forest, 220 m, 26.xi-24.xii.1983, M. Verhaagh [leg.], pitfall trap.

Oxyepoecus turgidus Kempf, 1969: 278. Holotype: 1W [Brazil], SP[São Paulo], Agudos, [Fazenda Santo Antônio], 23.ix.1954, [W.]W. Kempf [leg.]. Missing A1. Paratypes: 1W [Brazil], SP[São Paulo], Agudos, [Fazenda Santo Antônio], 25.i.1953, [W.]W. Kempf [leg., Coll. Kempf #740]. 2W [Brazil], GO[Goiás], Anápolis, [savanna near the Franciscan minor seminary], 18.iii.1964, [W.]W. Kempf [leg., Coll. Kempf #3852]. 1W [Brazil], GO[Goiás], Anápolis, [savanna near the Franciscan minor seminary], 2.i.1966, [W.]W. Kempf [leg., Coll. Kempf #4300]. Missing L2 tibia and tarsus. Junior synonym of O. inquilinus (Kusnezov, 1952a: 720) by Kempf, 1974a: 489.

Oxyepoecus vivax Kempf, 1974a: 508. Holotype: 1M [Brazil], MG[Minas Gerais], Pedra Azul, [15º59'S 41º07'W], 800 m, xi.1972, C.A.C. Seabra & M. Alvarenga [leg., Coll. Kempf] #8836. Missing A2/A3.

Genus Phacota Roger, 1862b: 260

Combination in Monomorium by Fernández, 2007b: 130.

Genus Solenopsis Westwood, 1840: 86

Labauchena acuminata Borgmeier, 1949: 208. Cotypes[Syntypes]: 3G [Argentina, Provincia] Salta, Colonia Juarez, [xii.1948], Luna [leg., Coll. Kusn.] #3272. One gyne missing A2; all gynes just with W4. Junior synonym of L. daguerrei Santschi, 1930: 81 by Borgmeier, 1959: 318; combination in Solenopsis by Ettershank, 1966: 140.

Solenopsis albidula var. postbrunnea Forel, 1913: 220. Cotypes[Syntypes]: 9W [Brazil], Est[ado de] S.P.[São Paulo], Franca, [ii.1911, E. Garbe leg., MZSP], #15952. One worker compressed dorso-ventrally; one worker missing head. 1W [Brazil], SP[São Paulo], Franca, [no date, no collector, Coll. Borgm.] #2701. Comment: The specimen #2701 labeled as cotype is not mentioned in the original description, so we do not consider it a type.

Solenopsis blumi Buren, 1972: 20. Paratypes: 24W Uruguay, Colonia Suiza, Ruta 1, km 119, 11.iii.1969, [M.S.] Blum, [C.] Crisci & [J.] Carbonell [leg.]. One worker missing A1 funiculus; one worker missing A1; one worker missing L5 trochanter, femur, tibia and tarsus; one worker missing head. Comment: The MZSP bears a pin with three specimens labeled as both holotype and paratype. As the original description says that holotype was deposited in the National Museum in Washington, D.C., we consider that the MZSP specimens are paratypes. Junior synnonym of S. quinquecuspis Forel, 1913: 224 by Trager, 1991: 185.

Solenopsis brevicornis var. petropolitana Borgmeier, 1928: 35. Cotypes[Syntypes]: 14W [Brazil, Rio de Janeiro], Petrópolis, ["im hohlen zweig (branch hollow), 22.ix.1924, types in Nationalmuseum"], #325.

Solenopsis bucki Kempf, 1973: 30. Holotype: 1W [Brazil], RS[Rio Grande do Sul], Erechim, [peneirada de humo], vii.1960, F. Plaumann [leg., Coll. Kempf] #8098.

Solenopsis clytemnestra var. leda Forel, 1913: 221. Cotypes[Syntypes]: 3W [Brazil], R[io de] Janeiro, Sapopemba, [no date, no collector], #2299. One worker with disarticulated postpetiole and gaster glued on paper triangle. 3W [Brazil, São Paulo], Esta[ção] A[lto] d[a] Serra[Linha de São Paulo/Santos, Serra do Mar (Lüderwaldt, 1918)], PP[Paranapiacaba, Santo André, viii.1910, E. Schwebel leg., MZSP] #17306. 8W [Brazil, São Paulo], A[lto] da Serra, [x.1910, E. Schwebel leg., Coll. Borgm.] #2121. Comment: Original labels of the specimens #17306 and #2121 were associated with new labels, which list "viii.1910". According the MZSP catalogue of identifier numbers the specimens #17306 were collected on viii.1910, and for the specimens #2121 the correct date is x.1910, according Thomaz Borgmeier catalogue.

Solenopsis clytemnestra var. strangulata Forel, 1913: 221. Cotypes[Syntypes]: 6W [Brazil], MG[Minas Gerais], Christina[Cristina, viii.1912, H. Lüderwaldt leg., MZSP], #17194. 12W [Brazil], MG[Minas Gerais], Cristina, [viii.1912, no collector, Coll. Borgm.] #2132. One worker missing head; one worker missing A1, A2 last segments of the funiculus, L3/L5/L6 trochanter, femur, tibia and tarsus, L4 last tarsus, treated with Barber's fluid.

Solenopsis corticalis var. margotae Forel, 1908: 364. Cotypes[Syntypes]: 5W 2G [Brazil, São Paulo, São Paulo], Ypiranga[Ipiranga, 8.x.1906], H. Lüderwaldt [leg., MZSP] #5588. 2W [Brazil, São Paulo, São Paulo], Ypiranga[Ipiranga, 8.x.1906], H. Lüderwaldt [leg., Coll. Borgm.] #2109. One worker missing L2 trochanter, femur, tibia and tarsus, treated with Barber's fluid.

Solenopsis decipiens abjecta var. ignobilis Forel, 1913: 220. Cotypes[Syntypes]: 6W [Brazil], Est[ado de] St[Santa] Cath[Catarina], Harmonia[Ibirama, MZSP, #15461. Unavailable name by Bolton, 1995: 388.

Solenopsis decipiens scelesta Forel, 1908: 364. Cotypes[Syntypes]: 3W [Brazil], Est[ado de] S[ão] P[aulo], Baurú, [9.xi.1906], B. Dreher col., [MZSP] #5982. 4W [Brazil, São Paulo, Baurú, 9,xi,1906, B. Dreher leg., Coll. Borgm.] #2127. One worker just with gaster; one worker missing A1, A2 last segment of the funiculus, L2/L3/L4/L5 trochanter, femur, tibia and tarsus, L6; both spesimens were treated with Barber's fluid.

Solenopsis franki Forel, 1908: 364. Syntypes: 8W [Brazil], SP[São Paulo], S[ão] Paulo, Ypiranga[Ipiranga], 16.ii.1906, J.[L.] Lima col., MP[MZSP] #2354. One worker missing A1/L1; one worker missing head, treated with Barber's fluid; one worker missing A1/A2, L1 tarsus, disarticulated L2 glued on the paper triangle, treated with Barber's fluid. Cotypes[Syntypes]: 4W [Brazil], SP[São Paulo], S[ão] Paulo, Ypiranga[Ipiranga], 16.ii.1906, J.[L.] Lima col., MP[MZSP] #2354. One worker just the postpetiole and gaster. Comment: Original labels of these specimens were associated with new labels, which say "galho seco, aroeira, cupinzeiro" (dry branch, aroeira, termite nest).

Solenopsis franki idae Forel, 1908: 365. Syntypes: 5W 1G, Cotypes[Syntypes]: 4W [Brazil, São Paulo, São Paulo], Ypiranga[Ipiranga, Mata do Governo], 8.x.1906, J.[L.] Lima col., MP[MZSP] #5578. Dealate gyne missing A1/A2, L1/L2, L3 tibia and tarsus, gaster with a hole in the first tergite, mesonoto with a hole, treated with Barber's fluid. Comment: Original labels of these specimens were associated with new labels, which say "em taquara" (in bamboo).

Solenopsis iheringi Forel, 1908: 362. Cotypes[Syntypes]: 8W [Brazil], SP[São Paulo], Esta[ção] Raiz da Serra, [3.xii.1906], H. Lüderwaldt col., MP[MZSP] #6778. One worker missing A1 funiculus.

Solenopsis invicta Buren, 1972: 9. Holotype: 1W Brazil, MT[Mato Grosso], Cuiabá, Aq. Res. Sta., 16.ii.1971, [W.] Willard & [R.] Willians [leg.]. Paratypes: 15W Brazil, MT[Mato Grosso], Cuiabá, Aq. Res. Sta., 16.ii.1971, [W.] Willard & [R.] Willians [leg.]. One worker missing gaster; one worker missing L3 tibia and tarsus; one worker missing A1 funiculus. 6W Brazil, MT[Mato Grosso], Cuiabá, Victorino's house, 17.ii.1971, [W.] Willard & [R.] Willians [leg.]. 6W Brazil, MT[Mato Grosso], Cuiabá, 18.ii.1971, [W.] Willard & [R.] Willians [leg.]. 3W Brazil, MT[Mato Grosso], Coxim, 9.iii.1971, W.W. Kempf [leg.]. 7W 1G 1M [United States of America], Ala[bama], 20 mi[les] W[est] of Mobile, 3.i.1970, W.F. Buren [leg.]. 13W 2G [United States of America], Alabama, Daufin[Dauphin] Isl[and], 3.i.1970, W.F. Buren [leg.]. Junior synonym of S. wagneri Santschi, 1916: 380 by Bolton, 1995: 388; reinstated over S. wagneri because of broad usage, according to the ICZN (Shattuck et al., 1999).

Solenopsis joergenseni Santschi, 1919: 42. Cotypes[Not types]: 3W [Brazil, Minas Gerais], Pirapora, [vi.1912, E. Garbe, Coll. Borgm.] #2282. Comment: The MZSP has six more specimens with the number #2282 but without cotype label. Furthermore, the original description was based on specimens collected in Provincia Formosa, Argentina. So, we do not consider them as types.

Solenopsis leptanilloides Santschi, 1925a: 159. Cotype[Syntype]: 1W Argentina, [Provincia Santa Fe], Rosario, [no date, Jose Hubrich leg., Coll. Borgm.] #3825.

Solenopsis nigella st. prevalens Santschi, 1923: 257. Cotypes[Syntypes]: 2W [Argentina], Buenos Aires, Sierra Ventana, [C.] Bruch [leg.], Coll. Borgm. #3824. Junior synonym of S. nigella gensterblumi Forel, 1901: 298 by Creighton, 1930: 132.

Solenopsis photophila Santschi, 1923: 250. Cotype[Syntype]: 1W Argentina, Villaguay, [C.] Bruch [leg., Coll. Borgm.] #3829. Missing gaster.

Solenopsis pulleni Moreno & Mackay, 2001: 62. Paratype: 1W [United States of America], NM[New Mexico], Do[ñ]a Ana Co[County], Las Cruces, Jornada Playa, 8.x.1986, B. Pullen [leg.], #10. Comment: Unavailable name by Bolton, 2012.

Solenopsis reichenspergeri Santschi, 1923: 256. Cotype[Syntype]: 1W Brasil, Rio [de Janeiro, A.] Reichensperger, [Coll. Borgm.] #3833. Missing L3 trochanter, femur, tibia and tarsus.

Solenopsis succinea r. nicai Forel, 1913: 222. Cotypes[Not types]: 2W 3G 3M [Brazil], Est[ado de] S[ão] P[aulo], Franca, [ii.1911, E. Garbe leg., MZSP] #15898. One worker just with postepetiole and gaster. [Syntypes]: 6W, [Not types]: 2M [Brazil, São Paulo, Franca, xii.1910, E. Garbe, MZSP] #15892. One worker missing petiole and postpetiole; one worker missing A1; one male missing A1 last segment of the funiculus, L2/L3 trochanter, femur, tibia and tarsus. Cotypes[Syntypes]: 10W [Brazil], SP[São Paulo], Franca, [xii.1910, E.] Garbe [leg.], Coll. Borgm. #2123. One worker treated with Barber's fluid. Comment: The original paper describes only workers with the number #15892. Specimens from type series were sent to Borgmeier and received the number #2123. So the males #15892 and the specimens with the number #15898 are not types.

Solenopsis stricta st. foederata Santschi, 1923: 255. Cotypes[Syntypes]: 5W Brésil[Brazil, Santa Catarina], Blumenau, [no date, P.M.] Witte [leg.], Coll. Borgm. #3830. One worker just with postpetiole and gaster.

Solenopsis stricta st. foederata var. specularis Santschi, 1923: 256. Cotypes[Syntypes]: 2W Brésil[Brazil, Santa Catarina], Blumenau, [no date, P.M.] Witte [leg.], Coll. Borgm. #3831. One worker missing A1 funiculus; One worker missing A1 last segment of the funiculus, L3 trochanter, femur, tibia and tarsus. Unavailable name by Bolton, 1995: 391.

Solenopsis tridens Forel, 1911: 298. Syntypes: 3W [Brazil], BA[Bahia], Vila Nova["Vila nova = Bonfim" (Kempf, 1971), today named Senhor do Bonfim. "In the region of Caatingas" (Ihering & Ihering, 1911)], ii.1908, E. Garbe [leg.], MP[MZSP] #14211, Coll. Borgm. #2128. One worker missing head, L4 tibia and tarsus. Cotypes[Syntypes]: 6W [Brazil], Est[ado da] Bahia, Villa Nova[Senhor do Bonfim, ii.1908, E. Garbe leg.], MP[MZSP] #14211.

Solenopsis tridens var. substituta Santschi, 1925b: 236. Syntypes: 3W 3G [Brazil], SP[São Paulo], Pitangueiras, i.1925, [H. Lüderwaldt leg.], Coll. Borgm. #326. One worker missing head, L4 tibia and tarsus; dealate gyne; one gyne missing W2; one gyne missing W1/W3. Raised to species by Trager, 1991: 161. Comment: Santschi (1925b) referred to these specimens with the identifier number #20909, but according the MZSP catalogue of identifier numbers the specimen #20909 is a Chrysomelidae and the correct number for the syntypes is #20907. Besides that, specimens from type series were sent to Borgmeier and received the number #326.

Solenopsis (Diplorhoptrum) subterranea MacKay & Vinson, 1989, 175. Paratypes: 3W [United States of America], TX[Texas], Brazos Co[County], 10 km N[orth] Kurten, 5.v.[19]87, W. Mackay [leg.], #9149. One worker missing L3 trochanter, femur, tibia and tarsus. 6W [United States of America], TX[Texas], Brazos Co[County], 10 km N[orth] Kurten, 5.viii.1987, W. Mackay [leg.], #9636. One worker missing L1/L3 femur, tibia and tarsus, L2 last tarsus.

Carebara genus group

Genus Carebara Westwood, 1840: 86

Carebara concinna species complex

Carebara brevipilosa Fernández, 2004: 210. Paratype: 1S Brasil[Brazil], São Paulo, Agudos, 23.ii.1955[original publication says "1995"], W.W. Kempf [leg.], #1806. Comment: The correct year is 1955, since W.W. Kempf passed away in 1976 (Lopes, 1978).

Carebara escherichi species complex

Carebara angulata Fernández, 2004: 226. Paratypes: 2W Ecuador, Prov[incia] Napo, Limoncocha, 00º24'S 76º36'W, 20.vii.1973, C.W. Rettenmeyer [leg.], #5300. One worker missing A1/A2, L1/L2/L3 femur, tibia and tarsus.

Carebara audita Fernández, 2004: 230. Paratype: 1W Colombia, Nariño, [Orito], Territorio Kofán, bosque, 700 m, 28.ix.1998[original publication says "20.ix.1998"], E.L. González leg., winkler.

Carebara brasiliana Fernández, 2004: 227. Holotype: 1W Brasil, SP[São Paulo], Botucatu, [District] Rubião Jr[Júnior], folhiço, 19.xii.1990. B.H. Dietz col. Paratypes: 3W Brasil, SP[São Paulo], Salesópolis, E[stação] B[iológica de] B[oracéia], 12-17.vi.1997, B.H. Dietz & [C.I.] Yamamoto [leg.], winkler. One worker missing A1 funiculus, A2 last segment of the funiculus; one worker missing an antenna, disarticulated antenna and head glued on a paper triangle.

Carebara inca Fernández, 2004: 225. Paratype: 1W Peru, Madre de Dios, [Parque Nacional del Manu, Estación Biológica] Cocha Cashu, [in litter wet forest], 19º90'S 71º36W, 27x.1999, D.W. Davidson [leg.].

Carebara nuda Fernández, 2004: 231. Holotype: 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, xii.[19]57, Fritz Plaumann [leg.], xaxim. Missing L2/L3 trochanter, femur, tibia and tarsus, disarticulated L6 glued on the petiole. Paratypes: 8W Brazil, Nova Teutônia, 27º11'S 52º23'W, 300-500 m, 1.iv.1954[original publication says "xii.1957"], F. Plaumann [leg.]. One worker missing L1 tarsus; one worker missing A1; one worker missing L3/L6 trochanter, femur, tibia and tarsus; one worker missing L2/L3/L6 trochanter, femur, tibia and tarsus; one worker with disarticulated A1 and L2/L3/L5 glued on a paper triangle; one worker missing A2, L2 trochanter, femur, tibia and tarsus; one worker missing A2, L2 femur, tibia and tarsus; one worker missing L3 trochanter, femur, tibia and tarsus.

Carebara pilosa Fernández, 2004: 227. Holotype: 1W Brasilien[Brazil, Santa Catarina], Nova Teutônia, 27º11B 52º23L, 300-500 m, i.1957, Fritz Plaumann [leg.], xaxim. Paratypes: 4W [Brazil], SC[Santa Catarina], N[ova] Teutônia, iii.1971, F. Plaumann [leg.], #7863.

Carebara reina Fernández, 2004: 228. Paratype: 1W Colombia, Valle del Cauca, PNN[Parque Nacional Natural] Farollones de Cali, [Embalse del] Alto Anchicayá, 650 m, 29.iv.2001 [original publication says "19-24.vi.2001"], S. Sarria [leg.], #1912, winkler.

Carebara striata Fernández, 2004: 228. Paratype: 1W Brazil, Pará, Belém, Utinga Forest Preserve, 1968, in [frog stomach of] Dendrobates quinquevittatus, P.A. Silverstone [leg.]. Missing A2 last segment of the funiculus.

Carebara lignata species complex

Carebara coeca Fernández, 2004: 219. Holotype: 1S, Paratypes: 1S 2W Brasil GO[Goiás], Jataí, UFG[Universidade Federal de Goiás], CCA[Centro de Ciências Agrárias], BR[Road] 364, km 192, ninho no solo, 10.i.2000, J. Diniz & C. Lásaro cols. Holotype missing L3 trochanter, femur, tibia and tarsus, petiole, postpetiole and gaster.

Carebara elongata Fernández, 2004: 213. Paratype: 1W Colombia, [Departament] Bolivar, PNN[Parque Nacional Natural, Santuario de Fauna y Flora] Los Colorados, Sector Venado, 320 m, 15-19.ix.2000, E. Delefeout [leg.], #1731, winkler. Comment: Original paper says "Paratypes: (...) same data as type (...)" and "Holotype worker: Colombia, Magdalena, Tayron National Park, Zaíno section, 11º12'01"N 74º02'01"W, 50 m, in malaise trap, 30.ago-20.sep.2000, R. Henríquez leg. M 624, deposited in IAvH[Insect Collection, Instituto Humboldt, Colombia]".

Carebara globularia Fernández, 2004: 214. Paratypes: 1W Colombia, Nariño, Territorio Kofán, bosque, 700 m, 28.ix.1998, E.L. González [leg., winkler]. 1W Colombia, Amazonas, 7 km NW[Northwest] Leticia, forest, litter, 20-25.ii[original publication says "iii"].1972, S. [Peck] & J. Peck [leg.], #250. 2W Ecuador, Prov[incia] Napo, Limoncocha, 00º24'S 76º36'W, 280 m, 13.viii.1973, Marian Rettenmeyer [leg.], #67. One worker missing L3 femur, tibia and tarsus.

Carebara minuta Fernández, 2004: 217. Holotype: 1W [Brazil], MT[Mato Grosso], Utiariti, Rio Papagaio, 10.xi.1966, [K.] Lenko & [F.S.] Pereira [leg.], #4489. Paratype: 1W Brazil, MT[Mato Grosso], Utiriati[Utiariti], Papagaio River, 10.xi.1996[1966, K.] Lenko [& F.S. Pereira leg.], #4489.

Carebara reticulata Fernández, 2004: 218. Paratype: 1W Colombia, [Departament] Quindío, Génova, Vda[Vereda] El Dorado, Fca[Bosque] San Isidro, 04º12'98"N 75º47'80"W, 1500 m, 31.x.1999, E.[L.] González [leg.], w[inkler]. Comment: Original publication says "J. Sosa leg.".

Carebara tenua Fernández, 2004: 224. Holotype: 1S, Paratype: 1S Brasil, MG[Minas Gerais], B[elo] Horizonte, 29.vii.1975, R.L. Araújo [leg.], #6217. Holotype missing L1/L3 trochanter, femur, tibia and tarsus; paratype missing L1/L2 trochanter, femur, tibia and tarsus.

Oligomyrmex longiceps Santschi, 1929: 295. Typus[Lectotype]: 1W, Typus[Paralectotypes]: 1W 1M [Argentina], Sierras de Córdoba[Provincia Córdoba], La Granja, Alta Gracia, 1924, C. Bruch leg., #1683. Lectotype missing L3 trochanter, femur, tibia and tarsus. Combination in Carebara, designation of the worker lectotype and male paralectotype by Fernández, 2004: 216. Comment: In the MZSP there is another worker labeled as paralectotype from the same data as the male paralectotype not mentioned by Fernández, 2004: 216.

Oligomyrmex longiceps Santschi, 1929: 295. Typus[Not types]: 2M [Argentina], Sierras de Córdoba[Provincia Córdoba], La Granja, Alta Gracia, 1924, C. Bruch leg., #1683. One male missing A1; one male only with postpetiole and gaster. Combination in Carebara by Fernández, 2004: 216. Comment: These specimens were designated as types by Santschi (1929), but Fernández, 2004: 216 did not list them as paralectotypes.

Oligomyrmex stenopterus Kusnezov, 1952b: 184. Cotype[Syntype]: 1G [Argentina], Misiones, [Puerto] Bemberg, iii.1945, [N.] Kusnezov [leg.], #746. Missing W3, L5 trochanter, femur, tibia and tarsus, L6 tarsus. Combination in Carebara by Fernández, 2004: 219.

Genus Tropidomyrmex Silva, Feitosa, Brandão & Diniz, 2009: 34

Tropiomyrmex elianae Silva, Feitosa, Brandão & Diniz, 2009: 39. Holotype: 1W, Paratypes: 3W 3G 5M Brasil, MG[Minas Gerais], Tiradentes, Serra de São José, 21º05'30"S 44º10'36", 1267 m, [in ground nest of Anoplotermes pacificus Mueller, 1873 (Isoptera, Apicotermitinae)], 12.iv.2007, E.M. Cancello & M.A. Menezes cols. One gyne and one male, both dealate, were coated with gold for SEM examination.

ACKNOWLEDGMENTS

This work was supported by the Conselho Nacional de Desenvolvimento Científico e Tecnológico/CNPq.

Aceito em: 12/07/2012

Impresso em: 30/06/2013

APPENDIX

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Publication Dates

• Publication in this collection
  19 July 2013
• Date of issue
  2013

History

• Received
  12 July 2012
• Accepted
  30 June 2013