CATALOGUE OF THE DOLICHODERINAE, FORMICINAE AND MARTIALINAE (HYMENOPTERA: FORMICIDAE) TYPES DEPOSITED AT THE MUSEU DE ZOOLOGIA DA UNIVERSIDADE DE SÃO PAULO, BRAZIL

ABSTRACT

This catalogue lists types of three ant subfamilies (Dolichoderinae, Formicinae and Martialinae) housed in the Formicidae Collection of the Hymenoptera Laboratory, Museu de Zoologia da Universidade de São Paulo (MZSP), Brazil. It provides label information, morphological condition of specimens, nomenclatural changes, and type status, following recommendations of the International Code of Zoological Nomenclature (ICZN). Here we present information on types of 101 nominal species, of which 96 are still valid. Eight species are only represented by holotypes, 27 species only by paratypes, seven by holotypes and paratypes, 56 species by syntypes, two species by paralectotypes and one species by a lectotype and paralectotypes. With this issue we complete the publication of information regarding all MZSP ant types, summing 4,741 specimens of 892 nomenclaturally valid species.

KEY-WORDS:
Types; Collection; MZSP; Ants

RESUMO

Este catálogo inclui os tipos de formigas de três subfamílias (Dolichoderinae, Formicinae and Martialinae) depositados na Coleção de Formicidae do Laboratório de Hymenoptera, Museu de Zoologia da Universidade de São Paulo (MZSP), Brasil. Apresentamos a informação de rótulos e sobre a condição morfológica dos espécimes, bem como as eventuais mudanças nomenclaturais e novos status taxonômicos seguindo as recomendações do Código Internacional de Nomenclatura Zoológica. Listamos os tipos de 101 espécies de formigas (sendo 96 atualmente válidas); oito espécies listadas são representadas somente por holótipos, 27 por parátipos, sete por holótipos e parátipos, 56 apenas por síntipos, duas espécies por paralectótipos e uma espécie pelo lectótipo e paralectótipos. O presente trabalho completa a série de publicações sobre os tipos de formigas da coleção do MZSP, que inclui 4.741 espécimens tipo de 892 espécies nomenclaturalmente válidas.

PALAVRAS-CHAVE:
Tipos; Coleção; MZSP; Formiga

INTRODUCTION

Natural history museum collections have been recognized as rich sources of data that can contribute to a wide range of studies (Burgman et al., 1995; Wheeler et al., 2004; Johnson et al., 2011) due to its role as repositories of inventories (Lister & Climate Change Research Group, 2011) and cradle of the description, nomenclature and classification of biodiversity on Earth. The Ant Collection of the Museu de Zoologia da Universidade de São Paulo (MZSP) is one of the leading collections of Neotropical ants in number of types, species and specimens, as well as geographic coverage (Brandão, 2000). The earliest records in the collection correspond to material collected by Hermann von Lüderwaldt and Hermann von Ihering at the beginning of the XX century (Klingenberg & Brandão, 2005). In 1977, the acquisition of the Walter Wolfgang Kempf and Thomaz Borgmeier collections has considerably increased the collection size (Brandão, 1991). Since then, collection curators, students and several collaborators and associates have added important material to it (Esteves et al., 2011), either from inventories or as a result of taxonomic studies.

The present catalogue is the eighth in a series of papers that attempted to disclose all type specimens deposited at the MZSP ant collection. For the past eleven years, several students, technicians and scientists have contributed to the publication of seven catalogues, which include the fungus-growing ants (Myrmicinae: former Attini) (Klingenberg & Brandão, 2005); poneromorphs (Amblyoponinae, Ectatomminae, Heteroponerinae, Ponerinae and Proceratiinae) (Scott-Santos et al., 2008); Pseudomyrmecinae (Brandão et al., 2010); dorylomorphs (Aenictinae, Ecitoninae, Cerapachyinae and Leptanilloidinae) (Esteves et al., 2011); Dacetini and Solenopsidini (Ulysséa & Brandão, 2013); Cephalotini (Prado & Brandão, 2013); and of 12 traditional Myrmicinae tribes (Adelomyrmecini, Basicerotini, Blepharidattini, Crematogastrini, Formicoxenini, Lenomyrmecini, Myrmicini, Phalacromyrmecini, Pheidolini, Stegomyrmecini, Stenammini and Tetramoriini) (Ulysséa et al., 2015). Altogether, we gathered information on 892 name-bearing types and 4,741 specimens (Table 1). This represents one of the best assemblages of ant types worldwide, and arguably the richest ant collection in South America. In general, preservation conditions of type specimens and specimens are excellent.

TABLE 1
Summary of the numbers presented at the MZSP ant types catalogues. Abbreviations meaning: N = number of individuals, SPP = species, H = holotype only, H + P = holotype and paratype, P = paratype only, S = syntype, L = lectotype, L + PL = lectotype and paralectotype, PL = paralectotype, NE = neotype.

Ant types	N	SPP	H	H + P	P	S	L	L + PL	PL	NE
I. Attini	588	105	4	12	13	74	0	2	0	0
II. poneromorphs	410	112	24	19	32	35	2	0	0	0
III. Pseudomyrmecinae	128	37	3	2	21	10	1	0	0	0
IV. dorylomorphs	584	141	13	17	43	67	0	0	1	0
V. Cephalotini	544	43	4	17	15	5	1	0	0	1
VI. Dacetini and Solenopsidini	926	141	23	37	42	31	1	4	3	0
VII. Other Myrmicinae	1038	212	12	28	102	65	0	5	0	0
VIII. Formicinae, Dolichoderinae and Martialinae	523	101	7	9	27	56	0	1	2	0
∑	4741	892	90	141	295	343	5	12	6	1

Our catalogue series aim to foster future studies on the Neotropical ant fauna and to provide evidence of the remarkable ant collection housed at the MZSP. It also complies with what is recommended by the International Code of Zoological Nomenclature, in regards to institutional responsibility of type depositories (recommendation 72F, ICZN, 1999): “Every institution in which name-bearing types are deposited should (…) make them accessible for study, and publish lists of name-bearing types in its possession or custody”.

Here, we provide label information and conservation status of Dolichoderinae, Formicinae and Martialinae type specimens deposited at the MZSP. We also present an index of all the valid names listed in the present catalogue and their respective original combination and synonyms. The complete information regarding all names listed here can be found in Bolton (2016).

Finally, the present publication shall be followed by updates, the first listing names and making available images of the ant types issued in this catalogue series, and every time the collections amasses a significant number of ant type specimens. Also, we will update the classification adopted along the series, in special regarding the Myrmicinae tribal arrangement, which changed significantly lately, but may change further soon.

MATERIAL AND METHODS

Type specimen information is recorded as follows: original name combination, author, publication year, page number, type category, number of specimens of each sex or caste, place of collection, date, collector, original depository, collection number or collection code, conservation status (i.e., presence of broken or missing body parts), and taxonomic history.

Original name combinations are listed in alphabetical order by species name within the genus, tribe and subfamily they are currently classified. The terminology used for name-bearing types follows the International Code of Zoological Nomenclature (ICZN, 1999).

For each type specimen, label information was completely transcribed, and also contrasted with data published in the original description - easily found on AntWiki and AntCat websites (www.antwiki.org and www.antcat.org, respectively) - and with records listed in Thomaz Borgmeier, Walter Wolfgang Kempf, and MZSP collection catalogues. All supplemental information retrived from these additional sources is presented within brackets; for the sake of clarity, further information gathered from other sources were also added.

The following abbreviations were used: Coll. = collection; Coll. Borgm. = Tomáz Borgmeier Collection; Coll. Bruch = Carlos Bruch Collection; Coll. Kempf = Walter Wolfgang Kempf Collection; MP = Museu Paulista, whose Zoological section became the present MZSP in 1939; W = worker; G = gyne (Feitosa & Brandão, 2008); M = male; col(s). = collector(s); leg. = legit.

Further, abbreviations were adopted to indicate the conservation status of a specimen as follows: A = antenna, A1 = left antenna, A2 = right antenna; W = wing, W1 = anterior left wing, W2 = posterior left wing, W3 = anterior right wing, W4 = posterior right wing; L = leg, L1, L2 and L3 represent respectively the left legs and L4, L5 and L6 the right legs.

RESULTS AND DISCUSSION

This catalogue presents information on 523 specimens (485 workers, 26 gynes, and 12 males), which are name-bearing types of 101 species (96 valid, one nomen nudum, and four unavailable names). Eight names are typified only by holotypes, 27 only by paratypes, seven by holotypes and paratypes, 56 by syntypes, two by paralectotypes, and one species by a lectopype and paralectotypes. In total, 17 genera and three subfamilies are represented by these types in the MZSP Ant Collection: Dolichoderinae - Anillidris, Azteca, Dolichoderus, Dorymyrmex, Forelius, Gracilidris, Leptomyrmex, Linepithema, Tapinoma and Technomyrmex; Formicinae - Acropyga, Brachymyrmex, Camponotus, Lasius, Myrmelachista and Nylanderia; and Martialinae - Martialis (^{Appendix 1}).

Subfamily Dolichoderinae Tribe Dolichoderini Genus Dolichoderus Lund, 1831 :130

Monacis andinaKempf, 1962:36. Holotype: 1W, Paratypes: 6W, PERU, C[entral] Pichita Caluga, 2,150 m, 18.ix.1960, W. Weyrauch [leg.], #1145. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus bidens var. bahianaSantschi, 1921b:101. Cotypes[Syntypes]: 3W, BRAZIL, Ba[hia], Ilhéus, iv.1919, E. Garbe col., #2182. One worker with disarticulated L3. Combination in Hypoclinea by Kempf, 1972:119; in Dolichoderus by Shattuck, 1992:77. Junior synonym of Dolichoderus bidens (Linnaeus, 1758) by Mackay, 1993:40.

Hypoclinea curvilobaLattke, 1987:259. Paratype: 1W, COLOMBIA, Chocó, Rio between S[an] Juan and Rio Baudó, 20.ii.1976, R.R. Wilkerson leg. Worker with disarticulated L3 and gaster. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus debilis var. rufescensMann, 1912:40. Cotypes[Syntypes]: 2W 1G, BRAZIL, [Pará], Amazonas, Madeira-Mamoré, Mann & Baker [legs.], RR. Co. Camp. 41. One worker missing gaster. Combination in Monacis by Kempf, 1959:252; in Dolichoderus by Shattuck, 1992:77. Raised to species by Kempf, 1959:252.

Hypoclinea epetreiaLattke, 1987:262. Paratypes: 4W, VENEZUELA, T[erritório] F[ederal] Amazonas dept., Río Negro, Río Baria, 140 m, 00°55’N/66°10’W, 03.viii.1984, L.J. Joly & T.J. Demarmela [leg.]. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus germaini subsp. garbeiForel, 1911:305. Syntypes: 11W, BRAZIL, Ba[hia], Joazeiro[Juazeiro], xi.1907, E. Garbe [leg.], MP 14206. One worker only with gaster; two workers missing A2. Combination in Hypoclinea by Kempf, 1972:119; in Dolichoderus by Shattuck, 1992:77. Junior synonym of Dolichoderus diversusEmery, 1894 by Mackay, 1993:50.

Dolichoderus gibbosus var. integraForel, 1911:306. Syntypes: 6W, [BRAZIL], Goiaz[Goiás, although the original publication says São Paulo state], Crixas, 1908, Dietz [leg.], MP 10869. 1W, [same location and date], Luderwaldt [leg.]. Worker with disarticulated head, mesosoma, gaster and legs, and missing A2. Combination in Hypoclinea by Kempf, 1972:120; in Dolichoderus by Shattuck, 1992:77. Subspecies of D. quadridenticulatus (Roger, 1862) by Shattuck, 1992:77. Junior synonym of D. quadridenticulatus by Mackay, 1993:86.

Monacis inpaiHarada, 1987:600. Paratype: 1W, BRAZIL, Am[azonas], Manaus, Z.F.3, km 24, Faz[enda] Esteio, Florestal, 18.iii.1983, A.Y. Harada col. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus lauraeMackay, 1993:67. Paratypes: 2W, COLOMBIA, Risavalda, mistrato, 980 m, 30.viii.[19]91, F. Fernandes [leg.], #16141.

Monacis lobicornisKempf, 1959:267. Holotype: 1W, Paratypes: 76W, [BRAZIL], S[anta] C[atarina], Blumenau, 12.x.1921, M. Witte [leg.], Coll. Borgm. Nr. 180, One worker only with gaster. Paratypes: 7W, [BRAZIL], Goiás, Anápolis, W. Kempf [leg.], 12.ii.1958, [Coll. Kempf Nr.] 2261. Combination in Dolichoderus by Shattuck, 1992:77.

Monacis omacanthaKempf, 1972:251. Holotype: 1W, Paratypes: 10W, [BRAZIL], Am[azonas], Manaus, Res[erva] Ducke, 04.vi.1969, INPA #15, #21, #6648, #6688. Paratypes: 7W, [same location], 20.v.1971, #4, #9, 6559, 6579. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus piceusMackay, 1993:81. Paratype: 1W, COLOMBIA, Valle [de Cauca], Represa Calima, 21.iii.1967, R. Root & W. Brown [legs.].

Monacis setosaKempf, 1959:267. Holotype: 1W, [BRAZIL], Pará, Cachoeira do Breu, J. Sampaio [leg.], x.[19]28. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus tristisMann, 1916:463. Syntype: 1W, BRAZIL, [Pará], Abunã, Rio Madeira, Mann and Baker [legs.]. Syntype: 1W, [same location], Madeira-Mamoré, R.R. Co. Camp 30 m, MCZ 46-9103. Worker missing L2 and L3. Combination in Monacis by Kempf, 1959:256; in Dolichoderus by Shattuck, 1992:77.

Monacis validusKempf, 1959:244. Holotype: 1W, Paratypes: 3W 1G, COSTA RICA, Zent, Limón, 20.i.1957, F. Lara E. [leg.], en cacao. Paratypes: 10W, [PANAMA], Río Chinillo, 06.iv.1913[original publication says 1923], W.M Wheeler [leg.]. Paratype: 1G, [PANAMA], Canal Zone 3, 06.iv.1923, #304, W.M Wheeler [leg.]. Combination in Dolichoderus by Shattuck, 1992:77.

Dolichoderus voraginosusMackay, 1993:106. Holotype: 1W, [BRAZIL], S[ão] P[aulo], Agudos, C. Gilberti [leg.], v.1959. Comment: We found at the MZSP ant collection specimens of Dolichoderus septemspinosus var. pallidipes [3W, British Guiana, Brartica distr. Kalecoon], and of Formica bispinosa var. lucidior [2W 2G, Panama, Los Sabana, 17.xi.1911, Wm. M. Wheeler leg.] labeled as “cotypes”, although these names have never been formally published. These specimens can not thus be treated as types.

Tribe Leptomyrmecini Genus Anillidris Santschi, 1936 a:414

Anillidris bruchiSantschi, 1936a:414. Typus[Syntypes]: 3W, ARG[ENTINA], Missiones Est., Exp. Loreto, 09.ix.[1]933[original publication says 1934], Dr. A.A. Oglobin [leg.]. Combination in Linepithema by Kempf, 1972:135; in Anillidris by Shattuck, 1994:3. Comment: The MZSP ant collection has one gyne and one male labeled as “cotypes”; as these specimens were not mentioned at the original publication we do not consider them as types.

Genus Azteca Forel, 1878 :384

Azteca aesopusForel, 1908:392. Cotypes[Syntypes]: 4W, [Coll. Borgm.] Nr. 2155. Comment: According to Borgmeier’s catalogue the label information is “BRAZIL, Espírito Santo, Porto Cachoeira, E. Garbe col., 01.xii.1905”, but the original publication says “São Paulo, v. Ihering col.”.

Azteca alfari var. mixtaForel, 1908:386. Cotypes[Paralectotypes]: 4W, [Coll. Borgm.] Nr. 2153. One worker missing L3 tarsi. Junior synonym of Azteca alfariEmery, 1893 by Longino, 1989:5. Comment: According to Borgmeier’s catalogue the label information is “BRAZIL, Rio de Janeiro, Rio Sapopemba, 31.i.1906, Wacket leg.”, but the original publication says “San Bernardino, Paraguay or São Paulo, Brazil”. Longino (1989) designated the paralectotypes without studying or mentioning the specimens deposited at the MZSP ant collection.

Azteca andreaeGuerrero, Delabie & Dejean, 2010:54. Paratype: 1G, FRENCH GUIANA, [Sinnamary], Paracou [Research Station, 05°22’39”N/52°57’35”W], 24.vii.2008, Dejean, A. et al. col., hand collecting. Dealated gyne missing A1. Comment: At the MZSP ant collection there is one worker labelled as paratype, that can not be considered as type as the original publication did not mention this specimen.

Azteca diabolicaGuerrero, Delabie & Dejean, 2010:56. Paratype: 1G, PANAMA: San Lorenzo forest, 09°16’47.58”N/79°58’29.94”W, Ibisca [project], 17.x.2003, J. Schmidl [leg.], fog[ging] J-2.

Azteca goeldiiForel, 1906:245. Cotype[Syntype]: 1W, [BRAZIL, Acre, Aldeia Indígena] Porto Alegre, [Rio] Purus, Huber [leg., Coll. Borgm.] Nr. 3901.

Azteca goeldii st. croceiscapaForel, 1915:361 Cotypes[Syntypes]: 4W, [BRAZIL, São Paulo, Franca, Garbe col., ii.2011, Coll. Borgm.] Nr. 2164. One worker missing A1 and A2 funiculus. Nomen nudum attributed to Emery (Bolton, 1995).

Azteca iheringiForel, 1915:359. Cotypes[Syntypes]: 4W, [BRAZIL, Minas Gerais, Pirapora, vi.1912, E. Garbe col., Coll. Borgm.] Nr. 2163. One worker missing A2, L1, L2 and L3; one worker missing A2.

Azteca meridaLongino, 1991:1589. Paratype: 1G, V[ENE]Z[UELA], E[sta]do Trujillo, 14 km ESE Bocon, 09°12’N/70°09’W, 1,500-1,850 m, 25.viii.1987, ex Cecropia palmatisecta, Longino nº 1860-s. Dealated gyne.

Azteca paraensis var. bondariBorgmeier, 1937:250. Cotypes[Syntypes]: 6W 1G, [BRAZIL, Bahia, G. Bondar leg., ii.1930, Coll. Borgm.] Nr. 5265. Cotypes[Syntypes]: 3W, [BRAZIL, Bahia, Água Preta, 1933, Borgmeier leg., Coll. Bondar] Nr. 1842. Dealated gyne; one worker missing L1 tarsi. Comment: Borgmeier (1937) treated part of the specimens as types and part as paratypes, and did not designate one specimen as holotype. We consider them all as syntypes.

Azteca snellingiGuerrero, Delabie & Dejean, 2010:61. Paratype: 1W, PANAMÁ, Panama City, Parque Metropolitano, 26.ii.2008, N.B. Espirito Santo & S.P. Ribeiro [legs.].

Azteca ulei var. gibbiferaForel, 1908:392. Cotypes[Syntypes]: 4W, [BRAZIL, São Paulo, Ilha Vitória, F. Gunther leg., iv.1906, Coll. Borgm.] Nr. 2150, [MP 2492]. Junior synonym of Azteca muelleriEmery, 1893 by Longino, 1991:1591.

Genus Dorymyrmex Mayr, 1866 :494

Dorymyrmex goeldii subsp. fumigatusForel, 1908:385. Syntypes: 6W, [BRAZIL, São Paulo], Ypiranga[Ipiranga, date illegible], Lima col. One worker missing L1 and L4; one worker missing gaster.

Dorymyrmex jheringiForel, 1912:39. Syntypes: 4W, [BRAZIL, São Paulo, Franca, E Garbe leg.], MP 15.822. One worker missing L3 and L4. Combination in Dorymyrmex (Conomyrma) by Forel, 1913:243; in D. (Ammomyrma) by Kusnezov, 1952:429; in Conomyrma by Kempf, 1972:79; in Dorymyrmex by Shattuck, 1992:85.

Genus Forelius Emery, 1888 :389

Forelius andinusKusnezov, 1957:15. Paratypes: 7W 2G, ARGENTINA, Prov[ince] Salta, S[an] Antonio de los Cobres, 3,800 m, N.K.[Nicolas Kusnezov leg.]. One worker missing part of A1 funiculus.

Forelius bahianusCuezzo, 2000:227. Holotype: 1W, Paratypes: 4W 1G, BRAZIL, Bahia, [Ilhéus], Olivença, [x.]1997. Gyne only with mesosoma; two workers missing gaster.

Forelius damianiGuerrero & Fernández, 2008:53. Paratype: 1W, COLOMBIA: Magdalena, Santa Marta, Sector Puerto Mosquito Sierra Nevada de Santa Marta, 11°10’23.6”N/74°10’45”W, 90 m, Pitfall, 24-26.ii.2007, D. Ramírez leg., bosque muy seco Tropical (bms-T). Worker missing A1.

Forelius maranhaoensisCuezzo, 2000:255. Holotype: 1W, BRAZIL, Maranhão, S[ão] Luís [do] Maranhão, 1982, Wanda M. Villioti col. Worker missing L2 and A1 funiculus. Paratype: 1W, BRAZIL, Bahia, S[an]ta Rita de Cássia, Riacho Veredão 13, 18.viii.1991, Brandão, C.R.F. col. Paratypes: 2W, BRAZIL, Bahia, Estrada Planaltino, M. Souza, 30.xi.1990, 16 hs, Brandão & Diniz [legs.]. Two workers missing gaster. Paratypes: 7W, BRAZIL, Goiás, Alvorada do Norte, Faz[enda] Matos, 08-12.vii.1991, Brandão, C.R.F. col., 22, isca sardinha, dia-solo; pitfall, estação 12; 19, isca sardinha, dia-vegetação.

Genus Gracilidris Wild & Cuezzo, 2006 :59

Gracilidris pomberoWild & Cuezzo, 2006:62. Paratype: 1W, PARAGUAY, Pte Hayes, 5 km SSE Pozo Colorado, 23°33’S/58°46’W, 140 m, 05.xii.2002. A.L. Wild [leg.], #AW 1766, Humid Chaco, grazed Copernicia Forest, Nest in soil.

Genus Leptomyrmex Mayr, 1862 :652

Leptomyrmex relictus Boudinot, Probst, Brandão, Feitosa & Ward, 2016:663. Holotype: 1W, BRAZIL, Goiás, Niquelândia, Serra da Mesa, 14°20’39.5”S/48°10’21.1”W, 16.xi.2013, Cerrado s.s.[stricto sensu], inside nest of Cornitermes silvestrii Emerson, 1949. Paratypes: 1W 2M, [same data and colony as holotype], R06 [one worker]; TC08[one worke] and DEO2013-030 [one male]. Two workers with disarticulated L5.

Genus Linepithema Mayr, 1866 :496

Linepithema anathemaWild, 2007:27. Holotype: 1W, BRAZIL, M[inas] G[erais], 2 km S. Monte Verde, 22°54’S/46°03’W, 1,900 m, 26.viii.1996, under stone in shrubland, Ward, P.S. [leg.], PSW#13155. Worker missing gaster.

Linepithema cryptobioticumWild, 2007:41. Paratype: 1W, PARAGUAY, Boquerón, Enciso, 21°12’S/61°40’W, M. Leponce & T. Delsinne [legs.], #7597, Dry Chaco in sifted litter.

Linepithema neotropicumWild, 2007:91. Paratypes: 1W 1G 1M, PARAGUAY, Canindeyú, Res[erva Natural del Bosque] Mbaracayú, Jejuimi, 170 m, 24°08’S/55°32’W, 25.xi.2002. A.L. Wild [leg.], #AW1718.

Linepithema pulexWild, 2007:105. Paratypes: 2W 1M, PARAGUAY, Canindeyú, Res[erva Natural del Bosque] Mbaracayú, Jejuimi, 170M, 24°08’S/55°32’W, 12.xi.2002, A.L. Wild [leg.], base of Philodendron, Humid subtropical tall forest, acc. no. AW1678, #AW1678.

Linepithema tsachilaWild, 2007:109. Paratypes: 2W 1M, ECUADOR, Pichincha, ENDESA forest res., 00°06’N/79°02’W, 700 m, 05.xii.2003, A.L. Wild [leg.], 2º growth forest nest in rotting center of live tree, #AW2212.

Iridomyrmex riograndensisBorgmeier, 1928b:64. Lectotype: 1W, Paralectotypes: 4W, BRAZIL, Rio Grande do Sul, N[eu] Würtemberg, x.1914, [E.] Garbe leg., MP 19.113. Syntypes[Paralectotypes]: 8W, [BRAZIL, Rio Grande do Sul], MP 19.118. Combination in Linepithema by Shattuck, 1992:16. Junior synonym of Linepithema humile (Mayr, 1868) by Wild, 2004:1207.

Iridomyrmex melleus succineusForel, 1908:396. Syntypes: 4W, [BRAZIL, São Paulo, Alto da Serra, H. Luderwaldt leg., MP] 5607. Combination in Linepithema by Shattuck, 1992:16. Junior synonym of Linepithema iniquum (Mayr, 1870:392) by Wild, 2007:68.

Tribe Tapinomini Genus Tapinoma Foerster, 1850 :43

Tapinoma amazonaeWheeler, 1934b:185. Types[Syntypes]: 2W, BRAZIL, Pará, Baker, gift of W.M. Wheeler, MCZ 13-19, 21372. One worker missing A1.

Genus Technomyrmex Mayr, 1872 :147

Tapinoma fulvumWheeler, 1934b:184. Cotypes[Syntypes]: 3W, [PANAMA], Barro Colo[rado], I.C.Z.; 25.vi.[19]24, nº 5-5-8, W.M. Wheeler [leg.]. Combination in Technomyrmex by Shattuck, 1992:161. Senior synonym of Technomyrmex fulvum subsp. sublucidum (Wheeler, 1934b):185 by Bolton, 2007:120.

Subfamily Formicinae Tribe Camponotini Genus Camponotus Mayr, 1861 :35

Camponotus abdominalis subsp. cupiensForel, 1908:410. Syntypes: 4W, [BRAZIL], SP[São Paulo], Camp.[Campos] do Jordão, xii-1905, H. Luederwaldt [leg.], Coll. Borgm. Nr. 2206. One worker missing A2. Junior synonym of C. atriceps (Smith, 1858) by Hashmi, 1973:82.

Camponotus (Myrmoturba) bonariensis st. garbeiSantschi, 1922:102. Syntypes: 3W, [BRAZIL, Paraná, Castro, vii.15, E. Garbe leg., Coll. Borgm. Nr.] 2241. 1W 1G, [BRAZIL, Paraná, Castro, vii.15, E. Garbe leg., Coll. Borgm. Nr.] 2796. One worker missing L6 3º to 5º tarsus [2796]. Combination in C. (Tanaemyrmex) by Emery, 1925:79.

Camponotus (Neomyrmamblys) compositorSantschi, 1922:111. Syntypes: 5W, [BRAZIL, São Paulo], Ilha Alcatrazes, SP, [H.] Lüderwaldt [leg.], MP 20202. 1W, [BRAZIL], SP[São Paulo], I.[Island] Alcatrazes, x-1920, [H.] Luederwaldt [leg.], MP 20202, Coll. Borgm. Nr. 2507. One worker missing A1 last antennomer; one worker missing A1 last three antennomers; one worker missing L2 femur to tarsi [2507]. Combination in C. (Myrmaphaenus) by Emery, 1925:155.

Camponotus (Hypercolobopsis) coptobregmaKempf, 1968:411. Paratypes: 4W, BRASIL, Amazonas, Manaus, [km 21of the Manaus-Itacoatiara road], viii.1962, K. Lenko - col., #2300. One worker just with head and collapsed gaster; one worker just with head, petiole and gaster; one worker missing A1 funiculus, A2 and L5 femur to tarsi; one worker missing A1 4º to 11º antennomer, A2 last 5 antennomers, L1 and L2 tarsi, and L3 2º to 5º tarsus.

Camponotus (Myrmobrachys) crassus st. amazonicusSantschi, 1922:122. Syntypes: 1W 1G, [BRAZIL], BA[Bahia], Ilhéus, iv-1919, [E.] Garbe [leg.], MP 19993, Coll. Borgm. Nr. 3230. Worker missing A1 last 5 antennomers and A2 funiculus. Misspelled amazonensis by Emery, 1925:163.

Camponotus (Myrmobrachys) crassus st. amazonicus var. chrysothrixSantschi, 1922:123. Syntypes: 1W, [BRAZIL], BA[Bahia], Juazeiro, xi-1913, [E.] Garbe [leg.], MP 18711, Coll. Borgm. Nr. 2797. 1W, [BRAZIL], BA[Bahia], Juazeiro, xi-1913, [E.] Garbe [leg.], MP 18711, Coll. Borgm. Nr. 2229. One worker missing L1 3º to 5º tarsus [2797]. Unavailable name, see Bolton, 1995:92.

Camponotus (Myrmobrachys) crassus var. delabiatusSantschi, 1925:247. Syntypes: 2W, BRAS.[BRAZIL], Para[Pará], Monte Alegre, coll. Reichensperger, Coll. Borgm. Nr. 550. One worker missing A2, femur to tarsi of all right legs; one worker missing tarsi of L3 and L6.

Camponotus (Myrmobrachys) crassus var. picticornisSantschi, 1936a:421. Syntype: 1W, ARG.[ARGENTINA], Missiones, Est. Exp. Loreto, [11.iv.1932], Dr. A.A. Oglobin [leg.], 1992, [nest in dry twig]. Junior synonym of C. crassusMayr, 1862 by Wild, 2007:47.

Camponotus dalmasiForel, 1899:145. Syntype: 1W, COLOMBIE[COLOMBIA], Sta[Santa] Marta, [MP] 3344, 1899, “nid de termites avec” Crematogaster. Specimen missing the last tarsi of L1, L2 and L3. Combination in C. (Myrmorhachis): Forel, 1914:274; in C. (Myrmocladoecus) by Emery, 1925:166; Wheeler, 1934a:424.

Camponotus (Myrmaphaenus) escherichiMenozzi, 1926:71. Syntype: 1W, [BRAZIL], SP[São Paulo], Mogi das Cruzes, 1926, Escherich [leg.], Coll. Borgm. Nr. 4257. Worker missing A1. Junior primary homonym of C. escherichiEmery, 1925:97. Replacement name: C. fallatusBolton, 1995:98.

Camponotus (Dendromyrmex) fabricii var. acomaForel, 1908:418. Syntypes: 4W, [BRAZIL, Espírito Santo, H. v. Ihering, Coll. Borgm. Nr.] 2217. Combination in Dendromyrmex:Emery, 1925:173. Junior synonym of C. nidulans (Smith, 1860) by Fernández, 2002:65.

Camponotus fastigatus subsp. vagulusForel, 1908:403. Syntypes: 3W, [BRAZIL], SP[São Paulo], Bauru, 09-xi-1906, [O.] Dreher, MP 5976, Coll. Borgm. Nr. 2215. 5W, [BRAZIL, São Paulo, Bauru, 09-xi-1906, O. Dreher, MP] 5976. One worker missing L5 femur to tarsi [2215]; one worker missing A1 [2215]; one worker missing head [2215]; one worker with disarticulated gaster glued on the same triangle [5976]; one worker represented just by gaster [5976]. Combination in C. (Myrmaphaenus) by Emery, 1925:155. Raised to species by Wild, 2007:50.

Camponotus fastigatus subsp. veraeForel, 1908:403. Syntypes: 3W, [BRAZIL], SP[São Paulo], Ilha S.[São] Sebastião, 1906, [Fr.] Günther, MP 5392, Coll. Borgm. Nr. 2209. 5W, [BRAZIL, São Paulo, Ilha São Sebastião, 1906, Fr. Günther, MP] 5392. One worker missing L3 tarsi; one worker missing L3 2º to 5º tarsus [2209]; one worker missing the last 5 antennomers of A1 and A2; one worker missing A1 and A2 last 7 antennomers. Combination in C. (Myrmaphaenus) by Emery, 1925:155.

Camponotus (Neomyrmamblys) genatusSantschi, 1922:114. Syntypes: 2W 1G 1M, [BRAZIL], São Paulo, Ypiranga[Ipiranga, H. Luederwaldt leg., MP] 18936. Combination in C. (Myrmaphaenus) by Emery, 1925:155. Comment: There is one pin labeled as syntype and as Coll. Borgm. Nr. 2866 which correspondent specimens are missing. According T. Borgmeier’s catalogue, they should be one gyne and one male.

Camponotus iheringiForel, 1908:412. Syntypes: 3W, [BRAZIL], SP[São Paulo], Bauru, 09-xi-1906, [O.] Dreher, MP 5969. 2W, [BRAZIL], SP[São Paulo], Bauru, 09-xi-1906, [O.] Dreher, MP 5969, Coll. Borgm. Nr. 2214. 1W, [BRAZIL], SP[São Paulo], Bauru, 09-xi-1906, [O.] Dreher, MP 5969, Coll. Borgm. Nr. 2850. 4W, [BRAZIL, São Paulo, São Paulo, Ipiranga, 08.x.1906, J. Lima leg., MP] 5572. One worker missing A2 last 5 antennomers [2214]. Combination in C. (Myrmamblys) by Forel, 1914:271; in C. (Myrmobrachys) by Emery, 1920:260. Misspelled as iheringi by Emery, 1925:165. Senior synonym of C. (Myrmobrachys) iheringi var. bajulusEmery, 1925 by Wild, 2007:47.

Camponotus (Neomyrmamblys) iridisSantschi, 1922:115. Syntypes: 3W, [BRAZIL], BA[Bahia], Ilhéus, [E.] Garbe [leg.], MP 20020. One worker missing A2. Combination in C. (Myrmaphaenus) by Emery, 1925:155.

Camponotus kauraSnelling & Torres, 1998:5. Paratypes: 3W, PUERTO RICO, Caño Gorda, nr Guánica, [0-20 m], 26 Oct 1991, [R.R.] Snelling, [J.A.] Torres and [M.] Canals, Coll. RRS 91-15, in mangrove thicket, ex dead wood of black mangrove Avicennia germinans. 2W 1G, PUERTO RICO, Caño Gorda, nr Guánica, [0-20 m], 26 Oct 1991, [R.R.] Snelling, [J.A.] Torres and [M.] Canals, RRS 91-17, in mangrove thicket, ex dead wood of black mangrove Avicennia germinans.

Camponotus (Myrmothrix) lenkoiKempf, 1960a:398. Paratypes: 2W, [BRAZIL], Amapá, Rio Amaparí, km 185, 09.vii.1959, John Lane col. One worker missing L6 tarsi; one pin missing specimen.

Camponotus (Pseudocolobopsis) luederwaldtiSantschi, 1922:118. Syntypes: 5W, [BRAZIL], SP[São Paulo], Ilha Alcatrazes, x-1920, [H.] Lüderwaldt [leg.], Coll. Borgm. Nr. 2492. One worker missing A1; one worker missing L5 femur to tarsi; one worker with disarticulated gaster glued on the same triangle. Junior synonym of C. macrocephalusEmery, 1894 by Kempf, 1968:408; of C. geralensisEmery, 1920 by Shattuck & McArthur, 1995:122.

Camponotus maritimusWard, 2005:10. Paratypes: 2W, USA [UNITED STATES OF AMERICA], CA[California], San Mateo Co.[County], Jasper Ridge, 37°24’N/122°14’W, 150 m, 03.iv.2004, Ward, P.S. #15202, under stone, oak woodland.

Camponotus maculatus subsp. guatemalensis var. scheffleriForel, 1908:407. Syntypes: 1W, [BRAZIL], SP[São Paulo], Campos de Jordão, [08.ii.06, H. Lüderwaldt leg.], MP 2407, Coll. Borgm. Nr. 2837, [nest in soil]. 2W 1G, [BRAZIL], SP[São Paulo], Campos de Jordão, [08.ii.06, H. Lüderwaldt leg.], MP 2407, Coll. Borgm. Nr. 2252, [nest in soil]. One worker with collapsed head, mesossoma and gaster [2252]; one worker missing L4 femur to tarsi [2252]; gyne missing A2 last 10 antennomers [2252]; one worker missing A2 last 8 antennomers and L1 tibia to tarsi. Unavailable name by Bolton, 1995:122. Comment: 4W and 2G with MP’s number [2407] but without type and locality labels. As the original publication, the MP’s and T. Borgmeier’s catalogues did not mention the number of specimen belonging to the type series, we do not consider these seven specimens as syntypes.

Camponotus maculatus subsp. spengleriForel, 1908:406. Syntypes: 2W, [BRAZIL], RJ[Rio de Janeiro], Campos Itatiaia, [16.iv.1906, H.] Lüderwaldt [leg.], MP 2697, [nest under stone]. 6W, [BRAZIL, Rio de Janeiro, Campos Itatiaia, 16.iv.1906, H. Lüderwaldt leg., MP] 2697, [nest under stone]. 4W, [BRAZIL], RJ[Rio de Janeiro], Campos Itatiaia, 16-iv-1906, Luderw.[H. Lüderwaldt leg.], MP 2697, Coll. Borgm. Nr. 2202. One worker missing A1 funiculus [2202]; one worker missing L4; one worker missing L6 trochanter to tarsi, petiole and gaster; one worker just with part of mesossoma, middle and posterior legs. Combination in C. (Tanaemyrmex) by Emery, 1925:82. Currently subspecies of C. picipes (Olivier, 1792) by Emery, 1920:233.

Camponotus melanoticus var. catharinaeSantschi, 1939:323. Syntypes: 2W, [BRAZIL], SC[Santa Catarina], Ibirama[ex-Hammonia], 1915, [H.] Luederwaldt [leg.], MP 18990, Coll. Borgm. Nr. 2836, [nest under bark]. Combination in C. (Tanaemyrmex) by Kempf, 1972:69.

Camponotus (Myrmoturba) melanoticus st. valeriusSantschi, 1922:99. Syntypes: 4W, [BRAZIL], RS[Rio Grande do Sul], N. Wuertenberg[Neu Württemberg, today Panambi], x-1914, E. Garbe [leg.], MP 19103, Coll. Borgm. Nr. 2237. 3W [BRAZIL], RS[Rio Grande do Sul], N. Wuertenberg[Neu Württemberg, today Panambi], x-1914, E. Garbe [leg.], MP 19107, Coll. Borgm. Nr. 2243. One worker missing L5 femur to tarsi [19103]; one worker missing L5 tibia to tarsi [19103]; one worker missing L3 tibia to tarsi [19103]; one worker missing L2 tibia to tarsi; one worker missing L6 2º to 5º tarsus. Combination in C. (Tanaemyrmex) by Emery, 1925:81.

Camponotus (Myrmobrachys) oglobliniSantschi, 1934:32. Syntype: 1W, ARG.[ARGENTINA], Missiones, Est. Exp. Loreto, [20.iv.33], Dr. A.A. Oglobin [leg.], 2030, [nest under bark]. Missing A1 last two antennomers. Junior synonym of C. musRoger, 1863 by Kempf, 1978:37.

Camponotus (Myrmobrachys) phytophilusWheeler, 1934b:207. Syntypes: 4W, [MEXICO, Morelos], Cuernavaca, 22.vi.29, Till. Circinnata.[in Tillandsia cincinnata], gift of M. Wheeler, MCZ 12-15, 21600. One worker missing A1 and L3 tarsi.

Camponotus (Tanaemyrmex) polymorphicusMackay, López-Castro & Fernández, 2002:422. Paratypes: 2W 1G, COLOMBIA, Cundinamarca, Mpio. Chochontá, 05°01’N/73°42’W, 2,700-2,800 m, viii-2000, Camilo Lopez Castro [leg.], #19757, nest in soil and in living frailejón. One gye missing A1 and A2 funiculus, petiole and gaster; one gyne missing A1, A2 last antennomers, W2, L4 femur to tarsi, L5 2º to 5º tarsus, and L6 trochanter to tarsi.

Camponotus punctulatus subsp. termitariusEmery, 1902:297. Syntype: 1W, [BRAZIL], Rio Grande do Sul, St[São] Leopoldo, [MP] 3981, [living in termite nest, especially Eutermes and Anoplotermes, rarely in Cornitermes]. Combination in C. (Myrmoturba) by Forel, 1913:249; in C. (Tanaemyrmex) by Emery, 1925:79. Raised to species by Wild, 2007:48.

Camponotus (Myrmoturba) punctulatus r. minutior var. mediorufaForel, 1913:249. Syntype: 1W, REP. ARGENTINA, Prov. San Luiz, [Coll.] C. Bruch. Unavailable name. Material referred to C. punctulatus cruentusSantschi, 1922 by Santschi, 1922:104. Comment: At MZSP Ant Collection there are three workers from the type locality [2W, REP. ARGENTINA, Prov. San Luiz, C. Bruch; and 1W, REP. ARGENTINA, Prov. Mendonza, 10.v.12, C. Bruch] but not labeled as types. We were not able to infer about the status of this material since Forel (1913) did not specify the number of type specimens.

Camponotus (Tanaemyrmex) reburrus Mackay, in Mackay & Barriga, 2012:2. Paratypes: 1W 1M, ECUADOR, Orellana, Yasuni Research Station, 00°40’S/76°23’W, 200 m, 19-i-2009, P. Barriga and G. Alvia [leg.], #2009-021, nest in internodes of Cecropia. Male missing last two antennomers.

Camponotus (Myrmosphincta) reinaldiKempf, 1960b:456. Holotype: 1W, [BRAZIL], SP[São Paulo], Agudos, v-1958, R. Mueller [leg., Coll. Kempf] 2505. Paratype: 1W, [BRAZIL], SP[São Paulo], Agudos, v-1958, R. Mueller [leg., Coll. Kempf] 2505. Worker missing L3 tarsi. Comment: The holotype is a soldier (major worker) and the paratype a minor worker, both are in the same pin (Kempf, 1960b); has just labeled the holotype.

Camponotus (Myrmobrachys) senex subsp. textor var. obscuripesBorgmeier, 1929:214. Syntypes: 5W, [BRAZIL], Rio de Janeiro, GB[Guanabara, today Rio de Janeiro], Manguinhos, 30-xii-1922, A. Lutz [leg.], Coll. Borgm. Nr. 413. One worker missing L6 tarsi; one worker missing A2 last 4 antennomers and L6 tarsi; one worker missing A1 last 8 antennomers, A2 funiculus, L1 femur to tarsi, L5 4º and 5º tarsus. Unavailable name, see Bolton, 1995:114.

Camponotus (Pseudocolobopsis) subtruncatusBorgmeier, 1929:212. Lectotype[Syntype]: 1W, [BRAZIL], RJ[Rio de Janeiro], Petrópolis, xi-1918, T. Borgmeier [leg.], Coll. Borgm. Nr. 57, [in dry bamboo log]. Syntypes: 2W 1G, [BRAZIL], RJ[Rio de Janeiro], Petrópolis, xi-1918, T. Borgmeier [leg.], Coll. Borgm. Nr. 57. Gyne with some parts of body collapsed; one worker missing A1 last 5 antennomers and A2 funiculus; one worker missing L3 trochanter to tarsi; one worker missing L6 tibia and tarsi. Comment: William P. Mackay returned to the MZ one syntype labeled by him as lectotype dated 1998. Since this new type status was never published, we consider this specimen as part of the syntype series.

Camponotus (Neocolobopsis) scrobiferBorgmeier, 1928b:66. Syntypes: 2W, [BRAZIL], SP[São Paulo], Guarujá, 28-viii-1910, H. v. Jhering [leg.], Coll. Borgm. Nr. 3984, [in wood]. One worker with disaticulated head and part of A1 funiculus, both glued on the triangle; one worker missing A1, L2 tibia to tarsi, L3 femur to tarsi, L5 2º to 5º tarsus and L6 last tarsus. Junior synonym of C. coriolanusForel, 1912 by Kempf, 1968:410.

Camponotus (Tanaemyrmex) taniaeMackay & Delsinne, 2009. Paratypes: 2W, PARAGUAY, Boqueron, [Presidente Hayes, Parque Nacional Teniente] Enciso 2, [21.13747°S/61.50954°W, 24-25.ix.2003], Thibaut Delsinne [leg.], #11577, [dry chaco, pitfall trap].

Camponotus tainoSnelling & Torres, 1998:8. Paratypes: 3W, PUERTO RICO, El Verde [Field Station], Rio Grande, 200 m el., 25 July 1989, RRS#89-29b, [R.R.] Snelling & [J.A.] Torres Coll., “Tambonuco” rainforest, ex dead branch on tree. One worker missing A1 last three antennomers.

Camponotus trapezicepsForel, 1908:405. Syntypes: 2W, [BRAZIL], S.[São] Paulo, Ipiranga, [24.xi.05], J. Lima [leg., MP] 2250, [nesting in soil, 10-15 cm]. 5W 4M, [BRAZIL, São Paulo, Ipiranga, 24.xi.05, J. Lima leg., MP] 2250, [nesting in soil, 10-15 cm]. 3W, [BRAZIL], S.[São] Paulo, Ipiranga, [26].i-1907, Luderw.[H. Lüderwaldt leg.], MP 11239, Coll. Borgm. Nr. 2219. 3W 1G, [BRAZIL, São Paulo, Ipiranga, H. Lüderwaldt leg., MP] 7342. One worker missing A1 and A2; one worker missing A2; one male not completely emerged; one worker missing L3 2º to 5º tarsus [2219]; one worker missing A2 last 4 antennomers [7342]; gyne dealated missing the last 5 antennomers of A1 and A2 [7342]. Combination in C. (Myrmamblys) by Forel, 1914:272; in C. (Neomyrmamblys) by Santschi, 1921a:311; in C. (Myrmaphaenus) by Emery, 1925:156.

Camponotus (Myrmothrix) wheeleriMann, 1916:477. Syntype: 1W, [BRAZIL, Rondônia, ex Mato Grosso], Madeira-Mamore RR.[Railroad] Co. Camp 41, [306 km of Porto Velho], Rio Madeira, [W.M.] Mann & [F.] Baker [legs.], 9117. Combination in C. (Myrmoplatypus) by Santschi, 1921a:311.

Comment: The MZSP Ant Collection received from William P. Mackay a number of Camponotus specimens (listed below) labeled as holotype or paratype. These specimens are not considered types as the names have not been formally published thus far.

Camponotus acutus Mackay, 2003. Paratype: 1W, TX, Jeff Davis Co., Mclves Ranch, 30°37’06”N/104°01’03”W, 1,732 m, 02-vii-97, W. & E. Mackay, #17517, in branches.

Camponotus amblynodis Mackay, 1999. Paratype: 1W, COLOMBIA, Hulia, Rivera, 28 Dec. 83, W. & E. Mackay, #7118.

Camponotus angulophrysMackay, 2002. Paratypes: 2W 1G 1M, Kartabo, B.G., Jul. Aug. 1920, W.M. Wheeler, #444.

Camponotus caesariatus Mackay, 2003. Paratype: 1W, Avispas, PERU, Madre de Dios, 400 m, X-1/15-62, Pena.

Camponotus fernandezi Mackay, 2003. Paratype: 1W, COLOMBIA, Amazonas, Cagueta, Araracuara, x-94, G. Gangi Leg., GG327.

Camponotus fissurinatus Mackay, [no date]. Paratype: 1W, JAMAICA, St. James, Porto Elelfo, 18°28’N/77°53’W, 75 m, 11-iii-1984, J. Longino, #1120.

Camponotus flavobregmus Mackay, 2003. Paratype: 1W, COLOMBIA.

Camponotus gigantus Mackay, [no date]. Paratype: 1W, PANAMA, Bocas del Toro, 3 km W. Cont. Div. at Fortuna Hwy., el 1,000 m, 15 VI 1994, A.R. Gillogly.

Camponotus ithypilisMackay, 2002. Paratypes: 1W 1G, MEX, Hgo., 2 km W Orizatlán, 245 m, 08 Jun.1988, W. Mackay, #1091, nest in hollow twig.

Camponotus mariae Mackay, 2003. Paratype: 1W, DOMINICAN REPUBLIC, Botanical Garden, Mountain Isabel de Torres nr Puerto Plata, 28-v-1995, M. de Andrade.

Camponotus platyclyaens[?] Mackay, [no date]. Paratypes: 2W, COLOMBIA, Hulia, 1 km NE Rivera, 7 July 85, W. & E. Mackay, #7403.

Camponotus rubrigenysMackay, 2002. Paratypes: 2W, M. Bates, Bonacca ls., Hond., iv-35.

Camponotus virgatus Mackay, 1998. Holotype: 1W, Paratypes: 4W 1G, BRASIL, MG do Sul, 12 KS Areaga, 16 Oct 1989, W.P. Mackay, #12571, nest in twig.

Camponotus wettereri Mackay, 2003. Paratype: 1W, CA, Santa Cruz, 22-v-98, J. Wetterer, MB4.

Camponotus yanoviaki Mackay, 2004. Paratypes: 2W, PERU, Loreto, 28 km S Iquitos, 03.75°S/73.25°W, 18-vi-2004, S. Yanoviak, from hollow twig primary White sand forest.

Tribe Lasiini Genus Lasius ( Fabricius, 1804 :415)

Acanthomyops bureniWing, 1968:135. Syntypes: 6W, [UNITED STATES OF AMERICA], Wis.[Wisconsin, Barron County], Comstock, Aug.16.1941, Wm. F. Buren [leg.]. One worker missing A1 last two segments of the funiculus, L1, L3 and L6 tibia and tarsi, L5 2º to 5º tarsus; one worker missing L5 tibia and tarsi; one worker missing L5 and L6 tarsi. Combination in Lasius by Ward, 2005:13.

Lasius (Acanthomyops) pubescensBuren, 1942:405. Paratypes: 7W, [UNITED STATES OF AMERICA], Min.[Minnesota], Jenkins, Aug.11.1941, Wm. Buren [leg., taked from a sandy low mound nest in open woodlands]. One worker missing L1 and L5 2º to 5º tarsus. Combination in Acanthomyops by Creighton, 1950:433; in Lasius by Ward, 2005:13.

Lasius xerophilusMackay & Mackay, 1994:37. Paratypes: 2W 1M, [UNITED STATES OF AMERICA], N.W.[New Mexico], Otero Co.[County], White Sands Nat.[National] Mon.[Monument], 25 July 1992, W. Mackay [leg.], #16042, nest in soil, small mound > 10 cm. Male missing L2 tarsi.

Genus Nylanderia Emery, 1906 :133

Paratrechina burgesiTrager, 1984:483. Holotype: 1W, PERU, Machupichu, 2,600-2,800 m, 28.Feb.-01.Mar.67, W.L. Brown [leg.], H-176. Combination in Nylanderia by LaPolla et al., 2010:127.

Paratrechina (Nylanderia) troglodytesWeber, 1934:58. Syntype: 1W, CUBA, near Casa Harvard, Soledad, Cienfuegos, Aug.30, 1933, N.A. Weber [leg.], 31. Junior synonym of P. myopsMann, 1920 by Brown, 1955:135. Combination in Nylanderia by LaPolla et al., 2010:127.

Tribe Myrmelachistini Genus Brachymyrmex Mayr, 1868 :163

Brachymyrmex brasiliensisOrtiz & Fernández, 2014:22. Holotype: 1W, BRAZIL, São Paulo, Ubatuba, Parque Estadual da Serra do Mar, Núcleo Santa Virgínia, 23°19’S/45°06’W, 870-1,000 m, 22 Apr 2005, col. M. Uehara, USNM 00757748. Comment: In the original publication, the holotype locality is mistakenly informed as “Brazil, Rio de Janeiro, Nova Friburgo, Fazenda Barreto, 22°9’40.4712”S/42°31’27.4866”W, 1,068 m, 11-12 Jun 2011, col. T.M.S. Mesquita”.

Brachymyrmex delabieiOrtiz & Fernández, 2014:24. Holotype: 1W, BRAZIL, São Paulo, Tapiraí, 24°01’55.5”S/47°27’56”W, 08-14 Jan 2001, col. R.R. Silva & Eberhardt, Transecto 1 Winkler 23, USNM ENT 00757718. Missing A1 last antennomers, L2 coxa to tarsi, L3 trochanter to tarsi.

Brachymyrmex feitosaiOrtiz & Fernández, 2014:27. Holotype: 1W, Paratypes: 2W, Floresta da Tijuca, D. Federal, Brasil, 16.xii.1959, C.A. Campos Seabra [leg.], Coleção Campos Seabra, USNM ENT 00757694. One paratype missing L1 femur to tarsi.

Brachymyrmex longicornis var. hemiopsSantschi, 1923:668. Syntypes: 2W, [BRAZIL], S.[São] Paulo, Ipiranga, 07-iii-1906, [H.] Luederwaldt [leg.], MP 2371, Coll. Borgm. Nr. 2785. 2W, [BRAZIL, São Paulo, Ipiranga, 07-iii-1906, H. Luederwaldt leg., MP] 2371. One worker missing L1 and L4 5º tarsus [2785]; one worker with collapsed body, missing A1 and A2 [2785]; one worker missing A1 last 5 antennomers, A2 last three antennomers, L1 and L4 5º tarsus; one worker with collapsed body.

Brachymyrmex (Bryscha) micromegas Emery, in Santschi, 1923:675. Paralectotypes: 2W, [BRAZIL], SP[São Paulo], S[São] Paulo, Ipiranga, [xii.12, J. Lima & H. Lüderwaldt leg.], MP 17326. 2W, [BRAZIL, São Paulo, São Paulo, Ipiranga, xii.12, J. Lima & H. Lüderwaldt leg.], MP 17326. One worker missing A1 last 5 antennomers, A2 funiculus, L4 2º to 4º tarsus [USNM ENT 00757824]; one worker missing A1, A2, L2 and L3 tibia and tarsi, L5 3º to 5º tarsus, L6 trochanter to tarsi [USNM ENT 00757825]; one worker missing A1 and L3 tarsi [USNM ENT 00757827]; one worker missing L3 4º and 5º tarsus [USNM ENT 00757826]. Comment: Three major workers and 11 minor workers from original type series [BRAZIL, São Paulo, São Paulo, Ipiranga, xii.12, J. Lima & H. Lüderwaldt leg., MP 17326] were not designated as paralectotypes by Ortiz & Fernández (2014), but according to the International Code of Zoological Nomenclature (1999) we also should consider them as paralectotypes.

Genus Myrmelachista Roger, 1863 :162

Myrmelachista arthuri var. brunneicepsForel, 1908:397. Syntypes: 1W, [BRAZIL], SP[São Paulo], S.[São] Paulo, Ipiranga, [Mata do Governo, 08.x.06, J.] Lima leg., MP 5587. 10W, [BRAZIL, São Paulo, São Paulo], Ypiranga[Ipiranga], H. Lüderwaldt [leg., MP] 5587. 4W, [BRAZIL], SP[São Paulo], S.[São] Paulo, M. d. Governo, 08-x-1906, MP 5587, Coll. Borgm. Nr. 2264. Two workers missing A1; one worker missing A2; one worker missing L1 tibia and tarsi; one worker missing L2 5º tarsus, L4 3º to 5º tarsus and L6 4º and 5º tarsus; one worker missing L3 5º tarsus, L5 femur to tarsi, L6 4º and 5º tarsus; one worker missing L1 tibia and tarsi. Comment: The species name o is misspelled in the syntypes labels as var. brunnea.

Myrmelachista (Decamera) catharinae var. petropolitanaSantschi, 1936b:204. Syntypes: 3W, [BRAZIL, Rio de Janeiro, Petrópolis, Borgmeier. Combination in M. (Hincksidris): Kempf, 1972:149. One worker missing A1 and A2 funiculus.

Myrmelachista guyanensisWheeler, 1934b:198. Syntypes: 3W, B.G. [BRITISH GUYANA, today GUYANA], Kartabo, Jul. Aug. 1920, W.M. Wheeler Collection, M.C.Z. Cotype 13-15 21420, [nesting in dead twigs]. One worker missing L1 and L2 femur to tarsi.

Myrmelachista reticulataBorgmeier, 1928a:37. Syntypes: 5W 3G, [BRAZIL, Rio Grande do Sul], N[Nova] Petrópolis, 14-i-1927, P. Buck S.J. [leg.], Coll. Borgm. Nr. 1406. Combination in M. (Hincksidris) by Kempf, 1972:150. One worker missing A1 funiculus; one worker only with gaster.

Myrmelachista (Decamera) schachovskoiKusnezov, 1951:359. Paratypes: 8W, [ARGENTINA], Neuquén, S.[San] Martin de los Andes, N. Kusnezov [leg.]. Combination in M. (Hincksidris) by Kempf, 1972:150. One worker missing A1.

Tribe Plagiolepidini Genus Acropyga Roger, 1862 :242

Acropyga paramaribensisBorgmeier, 1933:263. Syntypes: 3W 3G, [SURINAM], Paramaribo, G.H. Buenzli [leg., in coffee plantation]. Combination in A. (Rhizomyrma) by Kempf, 1972:17. Junior synonym of A. exsanguis (Wheeler, 1909) by LaPolla, 2004:47. One gyne missing W2; one gyne dealated missing A1 funiculus.

Acropyga (Rhizomyrma) pickeliBorgmeier, 1927:287. Syntypes: 3W 1G, [BRAZIL, Paraíba, Bananeiras, ix.1927, B. Pickel O.S.B. leg., Coll. Borgm. Nr.] 1732. Junior synonym of A. decedens (Mayr, 1887) by Costa Lima, 1931:7. Revived from synonymy by Borgmeier, 1932:238. Junior synonym of A. goeldiiForel, 1893 by LaPolla, 2004:50. Gyne dealated.

Subfamily Martialinae Genus Martialis Rabeling & Verhaagh, 2008 :14913

Martialis heurekaRabeling & Verhaagh, 2008:14913. Holotype: 1W, BRAZIL, AM[Amazonas], EMBRAPA, km 28, hwy [highway] AM010, 30 km Manaus, 02°53’S/59°59’W, elev. 40-50 m, C. Rabeling [leg.], ex leaf litter at dusk primary lowland rainforest, CR030509-01, CASENT 0106181. Worker missing funiculus and right legs.

ACKNOWLEDGMENTS

We are grateful to Roberto Guerrero for helping us to clear up the terminology of some Azteca species’ types; Antônio dos Santos Silva who helped us to clear up the terminology applied to some type specimens; Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP Proc. 2012/21309-7) for financial support to M.A. Ulysséa; Programa de Capacitação Institucional do CNPq (MPEG/MCTI, CNPq/PCI-DC 313168/2016-4) for financial support to Prado, L.P.; and the reviewers for providing comments and corrections on the manuscript. Carlos Roberto F. Brabdão is grateful to Programa de Capacitação em Taxonomia - PROTAX (Processo CNPQ 440574/2015-3; FAPESP 2016/50378-8). We extend our recognition to all students, technicians, researchers and colleagues that contributed to enrich our holding for more than 100 years of summed work.

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• WHEELER, W.M. 1934b. Neotropical ants collected by Dr. Elisabeth Skwarra and others. Bulletin of the Museum of Comparative Zoology at Harvard College, 77:157-240.
• WILD, A.L. 2004. Taxonomy and distribution of the Argentine ant, Linepithema humile (Hymenoptera: Formicidae). Annals of the Entomological Society of America, 97:1204-1215.
• WILD, A.L. 2007. Taxonomic revision of the ant genus Linepithema. University of California Publications in Entomology, 126:1-151.
• WILD, A.L. & CUEZZO, F. 2006. Rediscovery of a fossil dolichoderine ant lineage (Hymenoptera: Formicidae: Dolichoderinae) and a description of a new genus from South America. Zootaxa, 1142:57-68.
• WING, W.M. 1968. Taxonomic revision of the Nearctic genus Acanthomyops. Cornell University Agricultural Experiment Station Memoir, 405:1-173.

• 1
  Editor Responsável: Kelli dos Santos Ramos
• Publicado com o apoio financeiro do Programa de Apoio às Publicações Científicas Periódicas da USP
• 3
  Os periódicos Papéis Avulsos de Zoologia e Arquivos de Zoologia estão licenciados sob uma Licença CC-BY da Creative Commons.

Appendix 1

Currently valid names of the listed taxa, followed by their original names and synonyms (if applicable).

Subfamily Tribe Genus	Valid species name	Original name	Synonym
Dolichoderinae
Dolichoderini
Dolichoderus	D. andinus	Monacis andina
	D. bidens	D. bidens var. bahiana
	D. curvilobus	Hypoclinea curviloba
	D. diversus	D. germaini subsp. garbei
	D. epetreius	Hypoclinea epetreia
	D. inpai	Monacis inpai
	D. laurae
	D. lobicornis	Monacis lobicornis
	D. omacanthus	Monacis omacantha
	D. piceus
	D. quadridenticulatus	D. gibbosus var. integra
	D. rufescens	D. debilis var. rufescens
	D. setosus	Monacis setosa
	D. tristis
	D. validus	Monacis validus
	D. voraginosus
Leptomyrmecini
Anillidris	A. bruchi
Azteca	A. aesopus
	A. alfari	A. alfari var. mixta
	A. andreae
	A. diabolica
	A. goeldii
	A. iheringi
	A. merida
	A. muelleri	A. ulei var. gibbifera
	A. paraensis bondari
	A. snellingi
Nomen nudum		A. goeldii st. croceiscapa
Dorymyrmex	D. goeldii fumigatus
	D. jheringi
Forelius	F. andinus
	F. bahianus
	F. damiani
	F. maranhaoensis
Gracilidris	G. pombero
Leptomyrmex	L. relictus
Linepithema	L. anathema
	L. cryptobioticum
	L. humile	Iridomyrmex riograndensis
	L. iniquum	Iridomyrmex iniquum succineus
	L. neotropicum
	L. pulex
	L. tsachila
Tapinomini
Tapinoma	T. amazonae
Technomyrmex	T. fulvus
Formicinae
Camponotini
Camponotus	C. atriceps	C. abdominalis cupiens
	C. bonariensis garbei
	C. compositor
	C. coptobregma
	C. coriolanus	C. scrobifer
	C. crassus	C. crassus picticornis
	C. crassus amazonicus
	C. crassus delabiatus
	C. dalmasi
	C. fallatus	C. escherichi
	C. fastigatus verae
	C. genatus
	C. geralensis	C. luederwaldti	C. macrocephalus
	C. iheringi	C. iheringi	C. iheringi bajulus
	C. iridis
	C. kaura
	C. lenkoi
	C. maritimus
	C. melanoticus catharinae
	C. melanoticus valerius
	C. nidulans	C. fabricii acoma
	C. ogloblini
	C. phytophilus
	C. picipes spengleri	C. maculatus spengleri
	C. polymorphicus
	C. reburrus
	C. reinaldi
	C. subtruncatus
	C. taniae
	C. taino
	C. termitarius	C. punctulatus termitarius
	C. trapeziceps
	C. vagulus	C. fastigatus vagulus
	C. wheeleri
Unavailable name		C. crassus amazonicus chrysothrix
		C. maculatus guatemalensis scheffleri
		C. punctulatus r. minutior var. mediorufa
		C. senex textor obscuripes
Lasiini
Lasius	L. bureni	Acanthomyops bureni
	L. pubescens
	L. xerophilus
Nylanderia	N. burgesi	Paratrechina burgesi
	N. myops	P. troglodytes
Myrmelachistini
Brachymyrmex	B. brasiliensis
	B. delabiei
	B. feitosai
	B. longicornis hemiops
	B. micromegas
Myrmelachista	M. arthuri brunneiceps
	M. catharinae petropolitana
Plagiolepidini	M. guyanensis
Acropyga	A. exsanguis	A. paramaribensis
	A. goeldii	A. pickeli
Martialinae
Martialis	M. heureka

Publication Dates

• Publication in this collection
  2017

History

• Received
  19 Jan 2017
• Accepted
  13 June 2017