SciELO - Scientific Electronic Library Online

vol.9 issue3-4Morphology of the egg and immature stages of Zagloba beaumonti Casey (Coleoptera, Coccinellidae)Mammals described in the Poranduba maranhense of Frei Francisco dos Prazeres author indexsubject indexarticles search
Home Pagealphabetic serial listing  

Services on Demand




Related links


Revista Brasileira de Zoologia

Print version ISSN 0101-8175

Rev. Bras. Zool. vol.9 no.3-4 Curitiba  1992 

Sobre a provável parafilia de Centrotinae Amyot & Serville (Homoptera, Membracidae)


About the probable paraphyly of Centrotinae Amyot & Serville (Homoptera, Membracidae)



Antônio José Creão-Duarte

Departamento de Sistamática e Ecologia, Universidade Federal da Paraíba, 58059-900 João Pessoa, Paraíba, Brasil




An alternative proposition for the analysis of the diagnostic states of the pronotum in the Centrotinae and Membracinae (sensu Strümpel) is presented. In this analysis, a linear transformation series is proposed for this character, departing from the state found in Tolania Stal, 1858 changing to an intermediate state in Centrotinae, which is still treated as plesiomorphic when compared with the one found in Membracinae. The transformation series thus defined, clarifies the problems related to the monophyly of the Membracidae, which is herein supported by the state of pronotum previously used to define the Centrotinae. This later taxon is treated as paraphyletic. The monophyly of the Membracinae is unquestioned, but its sister group is still unknown. Finally, this analysis permitted the reinterpretation of several characters, previously treated as homoplasies, now as synapomorphies.

Key words: Phylogeny, Centrotinae, Membracidae, Homoptera



Texto completo disponível apenas em PDF.

Full text available only in PDF format.




AMYOT, C.J.B. & A. SERVILLE. 1843. Histoire naturelle des insectes Hémiptères, Deuxième partie. Homoptères, Homoptera Latr. Paris, Librairie Encylopédique de Roret, p. 455-676.         [ Links ]

BUCKTON, G.B. 1903. A Monograph of Membracidae. London, Lovell Reeve & Co. Limited, 296p.         [ Links ]

DEITZ, L.L. 1975. Classification of the higher categories of the New World treehoppers (Homoptera: Membracidae). North Carolina Agric. Exp. Sta. Tech. Bull. 225:1-177.         [ Links ]

FUNKHOUSER, W.D. 1951. In: WYISMAN, P. Genera Insectorum: Homoptera family Membracidae, fasc. 208. Bruxelles, Luis Desmet-Vertenuil, 383p.         [ Links ]

GODING, F.W. 1926. Classification of the Membracidae of America. Journ. New York Entomol. Soc. 34 (4): 295-317.         [ Links ]

STRÜMPEL, H. 1972. Contribution to the Phylogeny of Membracidae Rafinesque (Homoptera: Auchenorhyncha). Zool. Jahrb. Syst. 99 (3): 313-407.         [ Links ]



Recebido em 01.II.1992; aceito em 20.V.1993.

Creative Commons License All the contents of this journal, except where otherwise noted, is licensed under a Creative Commons Attribution License