Austin, George T; Mielke, Olaf H.H

doi:10.1590/S0101-81751997000400012

Resumo

Fifteen species of Aguna Williams, 1927 (Hesperiidae: Pyrginae) are known from near Cacaulàndia in central Rondônia, Brazil; nine represent undescribed species. These nine, plus four additional new species discovered among material from other areas, are named and described: Aguna latifascia, Aguna coeloides, Aguna nicolayi, Aguna latimacula, Aguna penicillata, Aguna spicata, Aguna longicauda, Aguna panama, Aguna spatulala, Aguna similis, Aguna mesodenlala, Aguna squamalba, Aguna parva. The species of Aguna are placed into ten species groups based upon the characters of the male and female genitalia. These groups are described and their included species are discussed. Aguna williamsi Hayward, 1935, new synonymy is considered synonymous with Eudamus glaphyrus Mabille, 1888, revised status. Goniurus hypozonius Plötz, 1880, new status is considered a subspecies of Eudamus aurunce Hewitson, 1867. Neotypes are designated for Goniurus cholus Plötz, 1880, Papilio coelus Stoll, [1781], Goniurus gideon Plötz, 1880, and Goniurus hypozonius Plötz, 1880. Lectotypes are designated for Eudamus asander Hewitson, 1867, Eudamus panthius Herrich-Schäffer, 1869, Eudamus scheha Plötz, 1882, Eudamus mega-clesMabille, 1888, Eudamus leucogramma Mabille, 1888, Hesperia metophis Latreille, [1824], Eudamus aurunce Hewitson, 1867, and Eudamus glaphyrus Mabille, 1888. Preliminary identification keys are provided.

Lepidoptera; Hesperiidae; Aguna; Neotropical

Hesperiidae of rondônia, Brazil: Aguna Williams (Pyrginae), with a partial revision and descriptions of new species from Panama, Ecuador, and Brazil¹ 1 Contribution number 900 of the Departamento de Zoologia, Universidade Federal do Paraná.

George T. Austin^I; Olaf H.H. Mielke^II

^INevada State Museum and Historical Society. 700 Twin Lakes Drive, Las Vegas, Nevada 89107, USA

^IIDepartamento de Zoologia, Universidade Federal do Paraná. Caixa Postal 19020, 81531-990 Curitiba, Paraná, Brazil. Fellow CNPq

ABSTRACT

Fifteen species of Aguna Williams, 1927 (Hesperiidae: Pyrginae) are known from near Cacaulàndia in central Rondônia, Brazil; nine represent undescribed species. These nine, plus four additional new species discovered among material from other areas, are named and described: Aguna latifascia, Aguna coeloides, Aguna nicolayi, Aguna latimacula, Aguna penicillata, Aguna spicata, Aguna longicauda, Aguna panama, Aguna spatulala, Aguna similis, Aguna mesodenlala, Aguna squamalba, Aguna parva. The species of Aguna are placed into ten species groups based upon the characters of the male and female genitalia. These groups are described and their included species are discussed. Aguna williamsi Hayward, 1935, new synonymy is considered synonymous with Eudamus glaphyrus Mabille, 1888, revised status. Goniurus hypozonius Plötz, 1880, new status is considered a subspecies of Eudamus aurunce Hewitson, 1867. Neotypes are designated for Goniurus cholus Plötz, 1880, Papilio coelus Stoll, [1781], Goniurus gideon Plötz, 1880, and Goniurus hypozonius Plötz, 1880. Lectotypes are designated for Eudamus asander Hewitson, 1867, Eudamus panthius Herrich-Schäffer, 1869, Eudamus scheha Plötz, 1882, Eudamus mega-clesMabille, 1888, Eudamus leucogramma Mabille, 1888, Hesperia metophis Latreille, [1824], Eudamus aurunce Hewitson, 1867, and Eudamus glaphyrus Mabille, 1888. Preliminary identification keys are provided.

Key words: Lepidoptera, Hesperiidae, Aguna, Neotropical

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Texto completo disponível apenas em PDF.

ACKNOWLEDGMENTS. We thank the curators and others at various museums for the loan of specimens: L.D. and J. Y. Miller, and S.R. Steinhauser (AME), F.H. Rindge (AMNH), P.R. Ackery (BMNH), J.E. Rawlins (CM), B.P. Harris (LACM), S. S. Borkin (Milwaukee Public Museum), W. Mey (ZMHB), and J.M. Bums (USNM). Comments on our study and additional loans were graciously provided by R.J. Borth, J.P. Brock, W. W. McGuire, J.D. Turner, and E. Welling. We thank the following for field assistance: G. Bongiolo, J.P. Brock, O. Gomes, J.D. Turner, and F. and A. West. S.R. Steinhauser provided an extensive review of the manuscript. A.D. Warren informed us of records for Colima, Mexico. T.C. Emmel has provided encouragement and support since inception of investigations in Rondônia. The Schmitz family at Fazenda Rancho Grande always makes field studies in Rondônia a comfortable and enjoyable experience. The Conselho Nacional de Desenvolvimento Científico e Tecnológico kindly issued the authorization permits from the Ministério da Ciência e Tecnologia for our studies in Rondônia in collaboration with EMBRAPA/CPAC.

Recebido em 22.XI.1996, aceito em 12.XI.1997

Alayo, P.D. & L.E. Hernández. 1987. Atlas de las mariposas diurnas de Cuba (Lepidoptera: Rhopalocera). La Habana, Editorial Científico-Técnica, 149p.
Austin, G.T.; J.P. Brock & O.H.H. Mielke. 1993. Ants, birds, and skippers. Trop. Lepid. 4 (suppl. 2): 1-11.
Austin, G.T. & K. Johnson. 1995. Theclinae of Rondônia, Brazil: Areas, with descriptions of three new species (Lepidoptera: Lycaenidae). Trop. Lepid. 6:31-39.
Bailowitz, R.A. & J.P. Brock. 1991. Butterflies of southeastern Arizona. Tucson, Sonora Arthropod Studies, 342p.
Biezanko, CM. & O.H.H. Mielke. 1973. Contribuição ao estudo faunístico dos Hesperiidae Americanos. IV Espécies do Rio Grande do Sul, Brasil, com notas taxonômicas e descrições de espécies novas (Lepidoptera). Acta Biol. Paranaense, Curitiba, 2: 51-102.
Bridges, CA. 1988. Catalogue of Hesperiidae (Lepidoptera: Rhopalocera). Urbana, Illinois, published by the author.
Brown, P.M. & B. Heineman. 1972. Jamaica and its butterflies. London, E.W. Classey, 478p.
Brown Jr., K.S. & O.H.H. Mielke. 1967. Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera (continued): Lycaenidae, Pieridae, Papilionidae, Hesperiidae. Jour. Lepid. Soc. 21: 145-168.
De Jong, R. 1983. Annotated list of the Hesperiidae (Lepidoptera) of Surinam, with descriptions of new taxa. Tijds. v. Ent. 126: 233-268.
De la Maza E., R.G. & D. Gutieérrez. 1992. Ropaloceres de Quintana Roo, su distribucion, origen y evolucion. Rev. Soc. Mex. Lepid. 15: 3-43.
De la Maza J., A. & R.G. de la Maza E. 1985. La fauna de mariposas de Boca del Chajul, Chiapas, Mexico (Rhopalocera). Parte I. Rev. Soc. Mex. Lepid. 9: 23-44.
De la Maza R., R. 1987. Mariposas Mexicanas. México, D.F., Fondo de Cultura Económica, 302p.
Ehrmann, G.A. 1907. New tropical American Hesperiidae. Canad. Ent. 39:317-323.
Emmel, T.C.. & G.T. Austin. 1990. The tropical rain forest butterfly fauna of Rondônia, Brazil: species diversity and conservation. Trop. Lepid. 1: 1-12.
Evans, W.H. 1952. A catalogue of the American Hesperiidae in the British Museum (Natural History) Part II: Pyrginae - section 1. London, British Museum (Natural History), 246p.
Freeman, H.A. 1976. New Hesperiidae records for Mexico. Jour. Lepid. Soc. 30:62-67.
Godman, F.C. 1907. Notes on the American species of Hesperiidae described by Plötz. Ann. & Mag. Nat. Hist. 20: 132-155.
Godman, F.C. & O. Salvin. 1887-1901. Biologia Centrali-Americana. Zoology: Insecta. Lepidoptera-Rhopalocera. London, Taylor and Francis, vol. 2,782p., vol. 3, 112pls.
Hayward, K.J. 1933. Lepidópteros argentinos. Familia Hesperidae. Revta Soc. Ent. Argent. 5: 149-188.
______. 1935. Hesperiidae (Lep.) Argentinos. Notas y adiciones con decripción de una nueva especie. Revta Soc. Ent. Argent. 7: 131-137.
Hayward, K.J. 1948. Insecta, Lepidoptera (Rhopalocera), familia Hesperiidarum, subfamiliae Pyrrhopyginarum et Pyrginarum. In: U.R. Descole (Ed.). Genera et species animalium argentinorum. Buenos Aires, Guillermo Kraft, 389p.
Hernández, L.R.; D.S. Smith; N. Davies & A. Areces-Mallea. 1994. The butterflies and vegetational zones of Ghanahacabibes National Park, Cuba. Bull. Allyn Mus. 139: 1-19.
Herrich-Schäffer, G.A.W. 1869. Prodromus Systematis Lepidopterorum. Versuch einer systematischen Anordung der Schmetterlinge. Corr.-bl. zool.-min. Ver. Regensburg 23: 184-204.
Hewitson, W.C. 1867. Descriptions of one hundred new species of Hesperiidae. London, John Van Voorst, 25p.
Hoffmann, C.C. 1941. Catálogo sistemático y zoogeográfíco de los lepidópteros mexicanos. Sequnda parte. Hesperioidea. Ann. Inst. Biol., UNAM 12:237-294.
HORN, W. & P.H.I. Kahle. 1935-1937. Über entomologische Sammlungen. I. Kapitel: Sammlungen welche ihren Eigentümer gewechselt haben (nebst Angaben über Ausbeuten von Expeditionen, Privat-Sammlern und Händlern). Ent. Beih. Berl.-Dahlem 2: 1-160,3: 161-296,4: 297-388.
Lamas, G.R.; K. Robbins & D.J. Harvey. 1991. A preliminary survey of the butterfly fauna of Pakitza, Parque Nacional del Manu, Peru, with an estimate of its species richness. Publ. Mus. Hist. Nat. UNMSM 40: 1-19.
Latreille, P.A. [1824]. In: P.A. Latreille & J.B. Godart. Encyclopédie Méthodique. Histoire naturelle des insectes. Veuve Agasse, Paris, 9: 711-793, 9 (2): 329-828.
Mabille, P. 1888. Diagnoses de Lépidoptères (Hespérides) nouveaux. Le Naturaliste 10: 98-99.
______. 1891. Description d'Hespérides nouvelles. Ann. Soc. Ent. Belg. 35:LIX-LXXXVIII, CVI-CXXI, CLXVIII-CLXXXVII.
______. 1903-1904. Lepidoptera Rhopalocera. Family Hesperidae [sic]. In: P. Wytsman. Genera Insectorum 17: 1-210.
Mabille, P. & E. Boullet. 1912. Essai de revision de la famille des Hespérides. An. Sc. nat., Zool., Paris (9) 16: 7-159.
Mabille, P. & Vuillot. 1891. Novitates Lepidopterologicae 3-6, p. 17-48.
McGuiRE, W. W. & M.A. Rickard. 1976. New Hesperiidae records for Texas and the United States. Jour. Lepid. Soc. 30: 5-11.
Meerman, J.C. & T. Boomsma. 1993. Checklist of the butterflies of the Shipstern Nature Reserve. Occ. Pap. Belize Nat. Hist. Soc. 2: 37-46.
Mielke, O.H.H. 1971. Contribuição ao estudo faunístico dos Hesperiidae Americanos - II. distribuição geográfica das espécies de Aguna Williams, 1927, com descrição de uma espécie nova e um novo sinônimo. Arq. Mus. Nac, Rio de Janeiro, 54: 203-208.
______. 1973. Contribução ao estudo faunístico dos Hesperiidae Americanos. III. Espécies coletadas em duas excursões ao Pará e Amapá, Brasil (Lepidoptera). Acta Biol. Paranaense, Curitiba, 2: 17-40.
______. 1980. Contribuição ao estudo faunístico dos Hesperiidae Americanos. V. Nota suplementar as espécies de Pyrrhopyginae e Pyrginae do Rio Grande do Sul, Brasil (Lepidoptera). Acta Biol. Paranaense, Curitiba, 8/9: 7-17.
Mielke, O.H.H. & M.M. Casagrande. 1992. Lepidoptera: Papilionoidea e Hesperioidea coletados na Ilha de Maracá, alto Alegre, Roraima, parte de projeto Maracá, com uma lista complementar de Hesperiidae de Roraima. Acta. Amaz. 21 (1991): 175-210.
Moschler, H. B. 1879. Neue exotische Hesperiidae. Verh. zool.-bot. Ges., Wien, 28: 203-230.
Plötz, C. 1880. Die Hesperiinen-Gattung Goniurus Hüb. und ihre Arten. Bull. Soc. Imp. Nat. Moscou 55 (2): 1-22.
______. 1882. Die Hesperiinen-Gattung Eudamus und ihre Arten. Stett. ent. Zeit. 42: 500-504, 43: 87-101.
Scott, LA. 1986. The butterflies of North America, a natural history and field guide. Stanford Univ. Press, Stanford, California, 583p.
Seitz, A. 1907-1924. Die Gross-Schmett. der Erde. Vol. 5. Die amerikanischen Tagfalter., Stuttgart, Alfred Kernen, 1139p.
SMITH, D.S. & L.R. Hernández. 1992. New subspecies of Pseudochrysops bornoi (Lycaenidae) and Saliana esperi (Hesperiidae) from Cuba, with a new island record and observations on other butterflies. Caribbean Jour. Sci. 28: 139-148.
Smith, D.S.; L.D. Miller & J.Y. Miller. 1994. The butterflies of the West Indies and south Florida. Oxford, Oxford Univ. Press, 264 pp.
Steinhauser, S.R. 1975. An annotated list of the Hesperiidae of El Salvador. Bull. Allyn Mus. 29: 1-34.
______. 1981. A revision of the proteus group of the genus Urbanus Hubner. Lepidoptera: Hesperiidae. Bull. Allyn Mus. 62: 1-41.
______. 1986. A review of the skippers of the narcosius group of species of the genus Astraptes Hubner (sensu Evans, 1952) and erection of a new genus. Lepidoptera: Hesperiidae Bull. Allyn Museum 104: 1-43.
Stoll, C. 1781. In: P. Cramer. De Uitlandsche Kapellen, voorkomende in de drie Waereld-Deelen Asia, Africa en America, by een verzameld en beschreeven. 4. Amsterdam, Baalde and Utrecht, Wild, 252p.
Tilden, J.W. 1971. Aguna claxon (Hesperiidae) new to the United States. Jour. Lepid. Soc. 25: 293.
Vargas F., I.; A. Luis-Martinez; J. Llorrente-Bousquets & A. Warren. 1996. Butterflies of the state of Jalisco, Mexico. Jour. Lep. Soc. 50: 97-138.
Williams Jr., R.C. 1926. Studies in the Neotropical Hesperiidae. Paper I. Trans. Amer. Ent. Soc. 52: 61-88.
______. 1927. Studies in the Neotropical Hesperioidea. Paper II. Trans. Amer. Ent. Soc. 53: 261-292.

1

Contribution number 900 of the Departamento de Zoologia, Universidade Federal do Paraná.

Datas de Publicação

Publicação nesta coleção
16 Jul 2009
Data do Fascículo
Dez 1997

Histórico

Aceito
12 Nov 1997
Recebido
22 Nov 1996

This work is licensed under a Creative Commons Attribution-NonCommercial 4.0 International License.

[1] Alayo, P.D. & L.E. Hernández. 1987. Atlas de las mariposas diurnas de Cuba (Lepidoptera: Rhopalocera). La Habana, Editorial Científico-Técnica, 149p.

[2] Austin, G.T.; J.P. Brock & O.H.H. Mielke. 1993. Ants, birds, and skippers. Trop. Lepid. 4 (suppl. 2): 1-11.

[3] Austin, G.T. & K. Johnson. 1995. Theclinae of Rondônia, Brazil: Areas, with descriptions of three new species (Lepidoptera: Lycaenidae). Trop. Lepid. 6:31-39.

[4] Bailowitz, R.A. & J.P. Brock. 1991. Butterflies of southeastern Arizona. Tucson, Sonora Arthropod Studies, 342p.

[5] Biezanko, CM. & O.H.H. Mielke. 1973. Contribuição ao estudo faunístico dos Hesperiidae Americanos. IV Espécies do Rio Grande do Sul, Brasil, com notas taxonômicas e descrições de espécies novas (Lepidoptera). Acta Biol. Paranaense, Curitiba, 2: 51-102.

[6] Bridges, CA. 1988. Catalogue of Hesperiidae (Lepidoptera: Rhopalocera). Urbana, Illinois, published by the author.

[7] Brown, P.M. & B. Heineman. 1972. Jamaica and its butterflies. London, E.W. Classey, 478p.

[8] Brown Jr., K.S. & O.H.H. Mielke. 1967. Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera (continued): Lycaenidae, Pieridae, Papilionidae, Hesperiidae. Jour. Lepid. Soc. 21: 145-168.

[9] De Jong, R. 1983. Annotated list of the Hesperiidae (Lepidoptera) of Surinam, with descriptions of new taxa. Tijds. v. Ent. 126: 233-268.

[10] De la Maza E., R.G. & D. Gutieérrez. 1992. Ropaloceres de Quintana Roo, su distribucion, origen y evolucion. Rev. Soc. Mex. Lepid. 15: 3-43.

[11] De la Maza J., A. & R.G. de la Maza E. 1985. La fauna de mariposas de Boca del Chajul, Chiapas, Mexico (Rhopalocera). Parte I. Rev. Soc. Mex. Lepid. 9: 23-44.

[12] De la Maza R., R. 1987. Mariposas Mexicanas. México, D.F., Fondo de Cultura Económica, 302p.

[13] Ehrmann, G.A. 1907. New tropical American Hesperiidae. Canad. Ent. 39:317-323.

[14] Emmel, T.C.. & G.T. Austin. 1990. The tropical rain forest butterfly fauna of Rondônia, Brazil: species diversity and conservation. Trop. Lepid. 1: 1-12.

[15] Evans, W.H. 1952. A catalogue of the American Hesperiidae in the British Museum (Natural History) Part II: Pyrginae - section 1. London, British Museum (Natural History), 246p.

[16] Freeman, H.A. 1976. New Hesperiidae records for Mexico. Jour. Lepid. Soc. 30:62-67.

[17] Godman, F.C. 1907. Notes on the American species of Hesperiidae described by Plötz. Ann. & Mag. Nat. Hist. 20: 132-155.

[18] Godman, F.C. & O. Salvin. 1887-1901. Biologia Centrali-Americana. Zoology: Insecta. Lepidoptera-Rhopalocera. London, Taylor and Francis, vol. 2,782p., vol. 3, 112pls.

[19] Hayward, K.J. 1933. Lepidópteros argentinos. Familia Hesperidae. Revta Soc. Ent. Argent. 5: 149-188.

[20] ______. 1935. Hesperiidae (Lep.) Argentinos. Notas y adiciones con decripción de una nueva especie. Revta Soc. Ent. Argent. 7: 131-137.

[21] Hayward, K.J. 1948. Insecta, Lepidoptera (Rhopalocera), familia Hesperiidarum, subfamiliae Pyrrhopyginarum et Pyrginarum. In: U.R. Descole (Ed.). Genera et species animalium argentinorum. Buenos Aires, Guillermo Kraft, 389p.

[22] Hernández, L.R.; D.S. Smith; N. Davies & A. Areces-Mallea. 1994. The butterflies and vegetational zones of Ghanahacabibes National Park, Cuba. Bull. Allyn Mus. 139: 1-19.

[23] Herrich-Schäffer, G.A.W. 1869. Prodromus Systematis Lepidopterorum. Versuch einer systematischen Anordung der Schmetterlinge. Corr.-bl. zool.-min. Ver. Regensburg 23: 184-204.

[24] Hewitson, W.C. 1867. Descriptions of one hundred new species of Hesperiidae. London, John Van Voorst, 25p.

[25] Hoffmann, C.C. 1941. Catálogo sistemático y zoogeográfíco de los lepidópteros mexicanos. Sequnda parte. Hesperioidea. Ann. Inst. Biol., UNAM 12:237-294.

[26] HORN, W. & P.H.I. Kahle. 1935-1937. Über entomologische Sammlungen. I. Kapitel: Sammlungen welche ihren Eigentümer gewechselt haben (nebst Angaben über Ausbeuten von Expeditionen, Privat-Sammlern und Händlern). Ent. Beih. Berl.-Dahlem 2: 1-160,3: 161-296,4: 297-388.

[27] Lamas, G.R.; K. Robbins & D.J. Harvey. 1991. A preliminary survey of the butterfly fauna of Pakitza, Parque Nacional del Manu, Peru, with an estimate of its species richness. Publ. Mus. Hist. Nat. UNMSM 40: 1-19.

[28] Latreille, P.A. [1824]. In: P.A. Latreille & J.B. Godart. Encyclopédie Méthodique. Histoire naturelle des insectes. Veuve Agasse, Paris, 9: 711-793, 9 (2): 329-828.

[29] Mabille, P. 1888. Diagnoses de Lépidoptères (Hespérides) nouveaux. Le Naturaliste 10: 98-99.

[30] ______. 1891. Description d'Hespérides nouvelles. Ann. Soc. Ent. Belg. 35:LIX-LXXXVIII, CVI-CXXI, CLXVIII-CLXXXVII.

[31] ______. 1903-1904. Lepidoptera Rhopalocera. Family Hesperidae [sic]. In: P. Wytsman. Genera Insectorum 17: 1-210.

[32] Mabille, P. & E. Boullet. 1912. Essai de revision de la famille des Hespérides. An. Sc. nat., Zool., Paris (9) 16: 7-159.

[33] Mabille, P. & Vuillot. 1891. Novitates Lepidopterologicae 3-6, p. 17-48.

[34] McGuiRE, W. W. & M.A. Rickard. 1976. New Hesperiidae records for Texas and the United States. Jour. Lepid. Soc. 30: 5-11.

[35] Meerman, J.C. & T. Boomsma. 1993. Checklist of the butterflies of the Shipstern Nature Reserve. Occ. Pap. Belize Nat. Hist. Soc. 2: 37-46.

[36] Mielke, O.H.H. 1971. Contribuição ao estudo faunístico dos Hesperiidae Americanos - II. distribuição geográfica das espécies de Aguna Williams, 1927, com descrição de uma espécie nova e um novo sinônimo. Arq. Mus. Nac, Rio de Janeiro, 54: 203-208.

[37] ______. 1973. Contribução ao estudo faunístico dos Hesperiidae Americanos. III. Espécies coletadas em duas excursões ao Pará e Amapá, Brasil (Lepidoptera). Acta Biol. Paranaense, Curitiba, 2: 17-40.

[38] ______. 1980. Contribuição ao estudo faunístico dos Hesperiidae Americanos. V. Nota suplementar as espécies de Pyrrhopyginae e Pyrginae do Rio Grande do Sul, Brasil (Lepidoptera). Acta Biol. Paranaense, Curitiba, 8/9: 7-17.

[39] Mielke, O.H.H. & M.M. Casagrande. 1992. Lepidoptera: Papilionoidea e Hesperioidea coletados na Ilha de Maracá, alto Alegre, Roraima, parte de projeto Maracá, com uma lista complementar de Hesperiidae de Roraima. Acta. Amaz. 21 (1991): 175-210.

[40] Moschler, H. B. 1879. Neue exotische Hesperiidae. Verh. zool.-bot. Ges., Wien, 28: 203-230.

[41] Plötz, C. 1880. Die Hesperiinen-Gattung Goniurus Hüb. und ihre Arten. Bull. Soc. Imp. Nat. Moscou 55 (2): 1-22.

[42] ______. 1882. Die Hesperiinen-Gattung Eudamus und ihre Arten. Stett. ent. Zeit. 42: 500-504, 43: 87-101.

[43] Scott, LA. 1986. The butterflies of North America, a natural history and field guide. Stanford Univ. Press, Stanford, California, 583p.

[44] Seitz, A. 1907-1924. Die Gross-Schmett. der Erde. Vol. 5. Die amerikanischen Tagfalter., Stuttgart, Alfred Kernen, 1139p.

[45] SMITH, D.S. & L.R. Hernández. 1992. New subspecies of Pseudochrysops bornoi (Lycaenidae) and Saliana esperi (Hesperiidae) from Cuba, with a new island record and observations on other butterflies. Caribbean Jour. Sci. 28: 139-148.

[46] Smith, D.S.; L.D. Miller & J.Y. Miller. 1994. The butterflies of the West Indies and south Florida. Oxford, Oxford Univ. Press, 264 pp.

[47] Steinhauser, S.R. 1975. An annotated list of the Hesperiidae of El Salvador. Bull. Allyn Mus. 29: 1-34.

[48] ______. 1981. A revision of the proteus group of the genus Urbanus Hubner. Lepidoptera: Hesperiidae. Bull. Allyn Mus. 62: 1-41.

[49] ______. 1986. A review of the skippers of the narcosius group of species of the genus Astraptes Hubner (sensu Evans, 1952) and erection of a new genus. Lepidoptera: Hesperiidae Bull. Allyn Museum 104: 1-43.

[50] Stoll, C. 1781. In: P. Cramer. De Uitlandsche Kapellen, voorkomende in de drie Waereld-Deelen Asia, Africa en America, by een verzameld en beschreeven. 4. Amsterdam, Baalde and Utrecht, Wild, 252p.

[51] Tilden, J.W. 1971. Aguna claxon (Hesperiidae) new to the United States. Jour. Lepid. Soc. 25: 293.

[52] Vargas F., I.; A. Luis-Martinez; J. Llorrente-Bousquets & A. Warren. 1996. Butterflies of the state of Jalisco, Mexico. Jour. Lep. Soc. 50: 97-138.

[53] Williams Jr., R.C. 1926. Studies in the Neotropical Hesperiidae. Paper I. Trans. Amer. Ent. Soc. 52: 61-88.

[54] ______. 1927. Studies in the Neotropical Hesperioidea. Paper II. Trans. Amer. Ent. Soc. 53: 261-292.

Brasil

Brasil

Hesperiidae of rondônia, Brazil: Aguna Williams (Pyrginae), with a partial revision and descriptions of new species from Panama, Ecuador, and Brazil

Resumo

Datas de Publicação

Histórico