Cossura yacy sp. nov. |
Conical, longer than wide |
Two pairs of nuchal organs at lateral margins of the peristomium |
With tentacles observed through dissection |
Annulate, arising from median region of the 3rd chaetiger |
all capillary chaetae with smooth base and distally spinulosa |
Three long anal cirri without intercirral processes |
This study |
Cossura abyssalis Hartman, 1967 |
Bluntly rounded in front, wider dorsally, about as long as wide |
Unknown |
Unknown |
Arising from median region of the 3rd chaetiger |
|
|
Hartman, 1967 |
Cossura aciculata (Wu & Chen, 1977) |
Bluntly conical with round tip |
Not seen |
Unknown |
Annulate, arising from median region of the 3rd chaetiger |
Fine spinulosa are present along the outer margin |
About 10 lobes, without appendages, except for one small bulb located mid-ventrally |
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Cossura alba Hartman, 1967 |
Triangular, longer than wide |
Unknown |
Unknown |
Filament from border between chaetigers 4 and 5 |
With dense fine spines |
Unknown |
Hartman, 1967 |
Cossura bansei
Hilbig, 1996 Hilbig, B. 1996. Family Cossuridae Day, 1963. In: Blake, J. A.; Hilbig, B. & Scott, P. H. eds. Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Santa Barbara, Museum of Natural History, p. 385-404.
|
Prostomium about as long as wide |
Unknown |
Unknown |
Arising from posterior border of chaetiger 3 |
Fine spines are present along the outer margin |
With three anal cirri |
Hilbig, 1996 Hilbig, B. 1996. Family Cossuridae Day, 1963. In: Blake, J. A.; Hilbig, B. & Scott, P. H. eds. Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Santa Barbara, Museum of Natural History, p. 385-404.
|
Cossura brunnea
Fauchald, 1972Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Los Angeles, Allan Hancock Foundation. 561p.
|
Conical, anteriorly rounded |
Unknown |
Unknown |
Cylindrical filament arising from the middorsal of the 3rd chaetiger |
Fine spines are present along the outer margin |
With three long anal cirri |
Fauchald, 1972Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Los Angeles, Allan Hancock Foundation. 561p.
|
Cossura candida
Hartman, 1955Hartman, O. 1955. Endemism in the North Pacific Ocean, with emphasis on the distribution of marine annelids, and descriptions of new or little known species. In: Allan Hancock Found ed. Essays. The Natural Sciences in Honor of Captain Allan Hancock on the occasion of his birthday July 26 1955. Los Angeles, University of Southern California Press, p. 39-60.
|
Conical lobe, a little longer than wide |
Unknown |
A soft, lobed pouch extending from a short cylindrical base and terminating distally in about 12 digitate, subequal lobes |
Cylindrical filament arising from the middorsal of the 3rd chaetiger |
Fine spines are present along the outer margin |
With three long anal cirri |
Hancock, 1955 |
Cossura chilensis Hartmann-Schröder, 1965 |
Triangular shaped with wide round anterior margin |
Unknown |
With 2-3 digitate lobes |
Originating on anterior median border of chaetiger 3 |
Fine spines are present along the outer margin |
With three long anal cirri |
Soto & Lemus, 2016Soto E. H. & Lemus, D. 2016. Nueva descripción de Cossura chilensis Hartmann- Schröder, 1965 (Polychaeta: Cossuridae), de la Bahía de Valparaíso, Chile central con notas de su ecología. Revista de Biología Marina y Oceanografía 51:609-620.
|
Cossura coasta
Kitamori, 1960Kitamori, R. 1960. Two new species of cirratulid and Nephtydae (Annelida: Polychaeta). Bulletin of the Japanese Society of Scientific Fisheries 26:1082-1085.
|
Clearly demarcated with a blunt depressed cone |
Unknown |
Unknown |
Arising at chaetiger 3 |
Unknown |
With three long anal cirri |
Pillai, 2001 |
Cossura consimilis
Read, 2000Read, G. B. 2000. Taxonomy and distribution of a new Cossura species (Annelida: Polychaeta: Cossuridae) from New Zealand. Proceedings of the Biological Society of Washington 113:1096-1110.
|
Blunt conical, with round tip |
Unknown |
Unknown |
Arising on anterior median border of chaetiger 3 |
Fine spines are present along the outer margin |
With three long anal cirri |
Read, 2000Read, G. B. 2000. Taxonomy and distribution of a new Cossura species (Annelida: Polychaeta: Cossuridae) from New Zealand. Proceedings of the Biological Society of Washington 113:1096-1110.
|
Cossura dayi
Hartman, 1976Hartman, O. 1976. Polychaetous annelids of the Indian Ocean including an account of species collected by members of the International Indian Ocean Expeditions, 1963-1964 and a catalogue and bibliography of the species from India. Journal of the Marine Biological Association of India 16:191-252
|
Conical |
Unknown |
Unknown |
Unknown |
Fine spines are present along the outer margin |
Unknown |
Hartman, 1976Hartman, O. 1976. Polychaetous annelids of the Indian Ocean including an account of species collected by members of the International Indian Ocean Expeditions, 1963-1964 and a catalogue and bibliography of the species from India. Journal of the Marine Biological Association of India 16:191-252
|
Cossura delta
Reish, 1958Reish, D. J. 1958. Description of a new species of Cossura (Annelida: Polychaeta) from the Mississippi Delta. Journal of the Washington Academy of Sciences 48:53-55.
|
Conical |
Two pairs of nuchal organs |
The base bears from 15 to 20 digitate lobes |
The single long, cylindrical tentacle originates from the middorsal of the 3rd chaetiger |
Fine spines are present distributed along the outer margin, according to the type of chaetae |
Characterized by possessing 6 to 10 digitate lobes on either side of the anus, with three long anal cirri |
Reish, 1958Reish, D. J. 1958. Description of a new species of Cossura (Annelida: Polychaeta) from the Mississippi Delta. Journal of the Washington Academy of Sciences 48:53-55.
|
Cossura dimorpha (Hartman, 1976Hartman, O. 1976. Polychaetous annelids of the Indian Ocean including an account of species collected by members of the International Indian Ocean Expeditions, 1963-1964 and a catalogue and bibliography of the species from India. Journal of the Marine Biological Association of India 16:191-252) |
Depressed, broadly rounded in front |
Visible nuchal organs |
Unknown |
Annulate, arising from median region of the 3rd chaetiger |
With dense fine spines |
Unknown |
Hartman, 1976Hartman, O. 1976. Polychaetous annelids of the Indian Ocean including an account of species collected by members of the International Indian Ocean Expeditions, 1963-1964 and a catalogue and bibliography of the species from India. Journal of the Marine Biological Association of India 16:191-252
|
Cossura flabelligera Zhadan, 2017 |
Conical, with rounded tip, about as long as wide |
Not seen |
Unknown |
Branchial filament attached to mid-length of chaetiger 3 |
With dense fine spines |
Pygidium not found |
Zhadan, 2017 |
Cossura ginesi
Liñero-Arana & Díaz-Díaz, 2010Liñero-Arana I. & Díaz-Díaz Ó. 2010. A new species of Cossuridae (Annelida: Polychaeta) from Venezuela. Interciência35:789-792.
|
Trapezoidal, dorsoventrally flattened, with well-developed anterior lateral horns |
A pair of nuchal organs as lateral ciliated furrows |
With approximately 20 ciliated digitated lobes |
Arising from posterior border of chaetiger 2 |
First one as curved capillary with long tips and rows of dense fine spines and smooth spine-like tip |
Perianal pad vertically cleft, with radial corrugations forming lobes, and with three long slender cirri |
Liñero-Arana & Díaz-Díaz, 2010Liñero-Arana I. & Díaz-Díaz Ó. 2010. A new species of Cossuridae (Annelida: Polychaeta) from Venezuela. Interciência35:789-792.
|
Cossura heterochaeta
Orensanz, 1976Orensanz, J. M. 1976. Los anélidos poliquetos de la província biogeográfica Argentina. IX. Poecilochaetidae y Cossuridae. Comunicaciones Zoológicas del Museo de Historia Natural de Montevideo 10:3-8.
|
Conical |
Unknown |
Unknown |
Filament from middorsal of chaetiger 2 |
With dense fine spines |
Unknown |
Orensanz, 1976Orensanz, J. M. 1976. Los anélidos poliquetos de la província biogeográfica Argentina. IX. Poecilochaetidae y Cossuridae. Comunicaciones Zoológicas del Museo de Historia Natural de Montevideo 10:3-8.
|
Cossura hutchingsae
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Trapezium-shaped with wide round anterior margin |
Unknown |
Unknown |
Filament from border between chaetigers 2 and 3 |
With dense fine spines |
Unknown |
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Cossura keablei
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Rectangular or trapezium-shaped, with round or almost straight anterior margin |
Unknown |
Unknown |
Branchial filament arising from chaetiger 3 |
With dense fine spines |
Unknown |
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Cossura laeviseta Hartmann-Schröder, 1962 |
Conical |
A pair of nuchal organs |
Unknown |
Branchial filament arising from anterior of chaetiger 3 |
Smooth |
With three short anal cirri |
Egremy-Valdez, 2016 |
Cossura longocirrata
Webster & Benedict, 1887Webster, H. E. & Benedict, J. E. 1887. The Annelida Chaetopoda, from Eastport, Maine. U.S. Commission Fish & Fish 22:707-758.
|
Conical |
Not seen |
Unknown |
Arising from posterior part of chaetiger 2 |
Unknown |
With three long anal cirri |
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Cossura pettiboneae (Ewing, 1987Ewing, R. M. 1987. Review of the genus Cossurella (Polychaeta: Cossuridae) including descriptions of two new species and a key to the species of the world. Bulletin of the Biological Society of Washington 7:3-10.) |
Longer than wide, broadly rounded anteriorly |
Pair nuchal slits at postero-lateral prostomial margin |
Unknown |
Inserting between chaetigers 3 and 4 |
Rows of dense fine spines |
Unknown |
Ewing, 1987Ewing, R. M. 1987. Review of the genus Cossurella (Polychaeta: Cossuridae) including descriptions of two new species and a key to the species of the world. Bulletin of the Biological Society of Washington 7:3-10.
|
Cossura platypus Zhadan, 2017 |
Triangular in lateral view, longer than wide |
Not seen |
with 10 buccal tentacles |
Filament attached to middle-length of chaetiger 3 |
With dense fine spines |
Divided by vertical cleft, with three short simple cirri, no intercirral process |
Zhadan, 2017 |
Cossura pseudakaina (Ewing, 1987Ewing, R. M. 1987. Review of the genus Cossurella (Polychaeta: Cossuridae) including descriptions of two new species and a key to the species of the world. Bulletin of the Biological Society of Washington 7:3-10.) |
Longer than wide, broadly rounded anteriorly |
Pair of nuchal slits at postero-lateral prostomial margin |
Unknown |
Filament attached to segmental border between chaetigers 2 and 3 |
With dense fine spines |
Rim slightly scalloped in appearance, simple ciliated, digitate caudal cirrus attached at apex of midventral notch |
Ewing, 1987Ewing, R. M. 1987. Review of the genus Cossurella (Polychaeta: Cossuridae) including descriptions of two new species and a key to the species of the world. Bulletin of the Biological Society of Washington 7:3-10.
|
Cossura pygodactylata Jones, 1956 |
Conical |
Unknown |
Unknown |
Arising from middle-posterior part of chaetiger 2 |
With dense fine spines |
With three long cirri and 8-12 shorter intercirral processes |
Jones, 1956 |
Cossura queenslandensis
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Blunt conical, with round tip and broadened base |
Unknown |
Unknown |
Filament attached to segmental border between chaetigers 2 and 3 |
With dense fine spines |
One or two cirri, in some cases they are short |
Zhadan, 2015Zhadan, A. 2015. Cossuridae (Annelida: Polychaeta: Sedentaria) from Australian and adjacent waters: the first faunistic survey. Records of the Australian Museum 67:1-24. http://dx.doi.org/10.3853/j.2201-4349.67.2015.1639
http://dx.doi.org/10.3853/j.2201-4349.67...
|
Cossura rostrata
Fauchald, 1972Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Los Angeles, Allan Hancock Foundation. 561p.
|
Bluntly conical |
Unknown |
Unknown |
Arising from chaetiger 3 |
With dense fine spines |
With three anal cirri |
Fauchald, 1972Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Los Angeles, Allan Hancock Foundation. 561p.
|
Cossura soyeri Laubier, 1964 |
Conical, anteriorly rounded |
Present |
Unknown |
Inserted at posterior margin of chaetiger 2 |
With dense fine spines |
With three anal cirri |
Parapar, 1995 |
Cossura sima
Fauchald, 1972Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Los Angeles, Allan Hancock Foundation. 561p.
|
Shorter than wide with a bluntly rounded anterior margin |
Unknown |
Unknown |
Arising from chaetiger 3 |
With dense fine spines |
With three anal cirri |
Fauchald, 1972Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Los Angeles, Allan Hancock Foundation. 561p.
|
Cossura modica
Fauchald & Hancock, 1981Fauchald, K. & Hancock, D. R. 1981. Deep-water polychaetes from a transect off central Oregon. Allan Hancock Monographs in Marine Biology 11:1-73
|
Conical |
Unknown |
Unknown |
Arising from chaetiger 4 |
all capillary chaetae have distal spinulosa |
|
Fauchald & Hancock (1981Fauchald, K. & Hancock, D. R. 1981. Deep-water polychaetes from a transect off central Oregon. Allan Hancock Monographs in Marine Biology 11:1-73) |