Miranda, María José

doi:10.11606/0031-1049.2016.56.10

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Articles • Pap. Avulsos Zool. 56 (10) • Dic 2016 • https://doi.org/10.11606/0031-1049.2016.56.10 copiar

Cladistic analysis of the Argentinian species of the genus Bostryx (Gastropoda, Stylommatophora) based on morphological evidence

Autoría SCIMAGO INSTITUTIONS RANKINGS

RESUMEN

El género Bostryx Troschel, 1847 es encontrado exclusivamente en Sudamérica, desde Ecuador hasta Chile, principalmente en relación a la cordillera de los Andes. La filogenia de Bostryx está basada en caracteres morfológicos de 24 especies de este género, 19 de las cuales correspoden a especies de Argentina, 5 son procedentes de Perú, además de 7 especies representantes de otros géneros, perenecientes a Bulimulidae y Odontostomidae. Una matriz de 31 taxa y 78 caracteres morfológicos (19 de los cuales son continuos y 59 son discretos) de conchilla, sistemas paleal y reproductor, fue analizada bajo pesos implicados con TNT. Las especies argentinas y peruanas del género se agruparon en un clado monofilético soportado por 14 caracteres de conchilla, los cuales son principalmente caracteres continuos de forma de la conchilla.

Palabras-Clave:
Bulimulidae; TNT; Pesos implicados; Caracteres continuos; Forma de la conchilla

Species	Dry Material	Alcohol Material
Bostryx birabenorum Weyrauch, 1965WEYRAUCH, W.K. 1965. Tres nuevos Gastrópodos terrestres de Sudamérica. Neotrópica, 11(35):71-76.	IFML 985a (1), 985 (2); MLP 11620 (3)	-
Bostryx catamarcanus (Parodiz, 1956)	MACN-In 17778 (1), 17778-1 (10); MLP 8061 (5), 11440 (2)	-
Bostryx cordillerae (Strobel, 1874)	IFML-Moll 15580; MACN-In 1354 (1), 1906 (1), 3234 (3), 9849 (9), 9850 (1), 9592 (5), 10001 (1), 12114 (1), 19625 (3), 32867 (1), 36944 (1); MCNL 361 (5); MLP 897 (4), 8062 (10), 9660, 9669 (8)	MACN-In 10001-2 (2)
Bostryx costellatus (Hylton Scott, 1971)	MACN-In 4221-1 (1), 4221-5 (1)	MACN-In 4221-1 (1)
Bostryx cuyanus (Pfeiffer, 1867)	IFML-Moll 15581 (4); MACN-In 8845 (1), 9592-3 (11), 10000 (4), 12113 (8); MCNL 1117 (10); MLP 10980 (1), 10981 (1), 41801 (2), s/n (6)	MACN-In 12113 (1), 30498 (1)
Bostryx famatinus (Doering, 1879)	MACN-In 3233 (1)	-
Bostryx martinezi (Hylton Scott, 1965)	IFML-Moll 15281 (5), 15608 (1), 15609 (1), 15610 (1), 15611 (1); MACN-In 25870 (2), 27208 (3), 27281 (2), 27431 (1), 36892 (1); MLP 10995 (1), 10996 (1), 10997 (2), 11011 (1), 11012 (1), 11445 (1), 11446 (3), 11447 (3), 11448 (2)	IBN 826 (1), 829 (1); IFML-Moll 15506 (1), 15507 (3), 15508 (1), 15509 (1), 16358 (2), 16360 (1), 16361 (1); MACN-In 27431 (1), 36892 (2)
Bostryx mendozanus (Strobel, 1874)	MACN-In 9850 (3); MLP 8082 (1)	-
Bostyx pastorei (Holmberg, 1912)	IFML-Moll 15573 (1), 15582 (1); MACN-In 1316 (6), 1517 (1), 1517-1 (4), 4916 (10), 9834 (5), 9917 (1), 9917-1 (10), 28800 (5); MLP 9671 (1)	IFML-Moll 15573 (1)
Bostryx peristomatus (Doering, 1879)	IFML-Moll 10881 (5), 15471 (11), 15595 (20); MACN-In 31619 (10); MLP 9512 (10), 9530 (3), 9671 (1)	IFML-Moll 15471 (3); MACN-In 10666 (1), 36934 (1), 37032 (1)
Bostryx reedi (Parodiz, 1947)	MACN-In 10001-1 (1), 9592-1 (7); MCNL 1116 (5)	MACN-In 9592 (1), 10001-1 (1)
Bostryx roselleus Miranda & Cuezzo, 2014MIRANDA, M.J. & CUEZZO, M.G. 2014. Taxonomic revision of the Bostryx stelzneri species complex, with description of a new species (Gastropoda: Orthalicoidea: Bulimulidae). American Malacological Bulletin, 32(1):1-20.	IFML-Moll 928 (15), 15487 (10), 15488 (10), 15489 (6), 15490 (6); MLP 11017 (1)	IFML-Moll 15487 (2), 15488 (4), 15489 (2), 15490 (10), 15491 (2), 15531 (1), 15534 (5), 15535 (2); MACN-In 39122 (7); MLP 13740 (7)
Bostryx rudisculptus (Parodiz, 1956)	MACN-In 380 (1), 380-1 (5); MLP 10171 (4)	IFML-Moll 16460 (2)
Bostryx scaber (Parodiz, 1948)	IFML-Moll 15472 (12), 15517 (3); MACN-In 3217 (1), 4221 (1), 4221-1 (5)	IFML-Moll 15472 (13)
Bostryx stelzneri (Dohrn, 1875)	IFML-Moll 548 (16), 1616 (14), 11019 (10); MACN-In 306 (10), 433 (1), 433-1 (10), 1018 (10), 6243 (1), 6243-1 (10), 7810 (4), 8838 (10), 9159 (2), 9229 (8), 9834 (1), 9852 (8), 10505 (3), 12335 (3), 14086-1 (6), 15711 (2), 17591 (1), 17591-1 (7), 17613 (5), 17777 (10), 17850 (4), 18356 (10), 19061 (1), 19061-1 (6), 19506 (10), 19619 (10), 21116 (10), 24305 (10), 24398 (7), 26645 (1); MLP 1298 (1), 1307 (2), 9527 (2), 9533 (5), 9534 (2), 9535 (4), 9536 (3), 9539 (2), 11291 (2),	IFML-Moll 14443 (2), 14446 (4), 15455 (3), 15456 (1), 15459 (1), 15461 (1), 15462 (1), 15464 (2), 15465 (3), 15467 (3), 15468 (3), 15469 (4), 15470 (3), 15473 (1), 15475 (4), 15477 (2), 15478 (3), 15479 (2), 15480 (3), 15482 (2), 15484 (3), 15485 (3), 15486 (3), 15492 (1), 15542 (2), 15543 (3), 15544 (5), 15545 (3), 16357 (2); MACN-In 1018 (1), 2466 (1), 24308 (1), 32861 (1)
Bostryx strobeli (Parodiz, 1956)	MACN-In 9916 (1), 9916-1 (14), 27283 (1); MLP 11009 (1), 11010 (1), 11018 (3)	MACN-In 9916 (1)
Bostryx torallyi (d’Orbigny, 1835)	IFML-Moll 122 (10), 160 (10), 15493 (10), 15522 (10), 15601 (10); MACN-In 1349 (1), 1349-1 (1), 3236 (4), 6969 (2), 8848 (1), 8848-1 (5), 19167 (10), 25987 (9), 32639 (1); MLP 9667 (4), 9676 (5), 9677 (2), 9678 (2), 9679 (2), 9681 (2), 9966 (3)	IFML-Moll 15493 (3), 15494 (3), 15495 (4), 15496 (3), 15498 (2), 15499 (2), 15500 (1), 15547 (1)
Bostryx tortoranus (Doering, 1879)	IFML-Moll 14877 (10), 15502 (8), 15505 (7), 15530; MACN-In 6516 (1), 6516-1 (4), 19461 (2), 19506 (1), 19590 (1), 27282 (4), 29450 (1), 30619 (1); 32686 (1); MLP 10995 (1), 10996 (1), 10997 (1), 13345 (1), 11443 (1), 11007 (1), 11149 (1), 11006 (1), 11442 (1), 11444 (1), 11014 (4), 11015 (1), 8072 (10)	IFML-Moll 15501 (2), 15502 (3), 15504 (1), 15505 (2), 15510 (1), 15574 (3), 16359 (1); MACN-In 30619 (1)
Bostryx willinki Weyrauch, 1964WEYRAUCH, W.K. 1964. Nuevos Gasterópodos terrestres y nuevos sinónimos de Sudamérica II. Acta Zoológica Lilloana, 20:33-60.	IFML-Moll 121 (1), 121 (3); MACN-In 19062 (1)	-
Bostryx solutus Troschel, 1847	IFML-Moll 1132 (20), 1133 (40), 15382 (8), 3060 (20), 3134 (20), 15384 (10), 15386 (30), 15387 (34), 16587 (7), 16639 (7)	-
Bostryx conspersus (Sowerby, 1833)	IFML-Moll 1390 (10), 3115 (6), 3348 (20), 3392 (13), 3410 (4), 12150 (10), 16331 (6), 16332 (7), 16354 (5), 16355 (10)	-
Bostryx rhodolarynx (Reeve, 1849)	IFML-Moll 10208 (7), 10613 (2), 14040 (15), 16524 (1), 16525 (12), 16526 (2), 16528 (3), 16546 (19)	-
Bostryx infundibulum (Pfeiffer, 1853)	IFML-Moll 14057 (18), 16430 (100), 16432 (150), 16435 (20)	-
Bostryx hamiltoni (Reeve, 1849)	IFML-Moll 12143 (100), 16434 (18), 16442 (10), 16551 (20)	-
Spixia tucumanensis (Parodiz, 1941)	IFML-Moll 522 (6), 802 (10), 14769 (12), 14770 (20), 15204 (10), 15353 (5)	IFML-Moll 14774 (10), 15350 (5)
Plagiodontes daedaleus (Deshayes, 1851)	IFML-Moll 772 (16), 1065 (15), 10884 (5), 14048 (20)	IFML-Moll 14203 (4)
Drymaeus poecilus (d’Orbigny, 1835)	IFML-Moll 177 (10), 828 (18), 14256 (4), 14265 (4), 14899 (1), 14900 (2), 15689 (3)	IBN 478 (2), IFML-Moll 15696 (2)
Bulimulus apodemetes (d’Orbigny, 1835)	IFML-Moll 13524 (2), 14922 (30), 14971 (25), 15624 (11)	IBN 491 (4)
Bulimulus sporadicus (d’Orbigny, 1835)	IFML-Moll 172 (8), 15652 (7), 15653 (21)	IBN 547 (2), 599 (2)
Bulimulus gracilis Hylton Scott, 1948	MLP 11014 (4), 11015 (1)	IFML-Moll 15655 (4)
Naesiotus munsterii (d’Orbigny, 1837)	IBN 187 (2)	IBN 187 (2)

Species	0	1	2	3	4	5	6	7	8	9	10	11	12	13	14	15	16	17	18
Bulimulus bonariensis	27.32-32.58	6.43-6.55	2.25-2.49	17.24-20.26	5.17-5.43	1.72-2.00	10.66-14.10	2.90-3.04	0.41-0.49	10.94-13.26	10.06-12.30	11.85-14.39	6.75-8.27	3.16-3.30	0.70-0.72	7.00-8.26	3.48-4.0	6.14-6.60	6.94-7.48
Bulimulus apodemetes	29.05-34.67	6.93-8.227	3.05-4.49	23.55-26.57	6.28-7.78	2.75-4.21	6.86-10.32	2.27-2.67	0.27-0.33	15.01-18.01	14.02-16.1	16.85-19.65	10.4-12.06	4.09-5.07	1.15-1.83	5.0-7.0	2.54-3.32	6.90-7.56	9.14-10.16
Bulimulus gracilis	29.88-32.30	6.56-7.18	2.27-2.55	21.85-23.31	5.53-5.83	1.93-2.11	8.95-10.89	2.42-2.88	0.36-0.44	12.06-12.60	11.12-11.84	15.02-16.36	8.18-8.74	3.47-3.49	0.77-0.81	5.94-6.22	3.45-4.19	6.16-7.15	7.51-9.55
Naesiotus munsterii	19.19-21.75	7.707	3.813	13.42-15.62	6.881	3.363	5.69-6.55	2.647	0.417	10.34-11.66	14.13	14.6	4.79-6.49	4.536	1.482	6.93-7.49	5.18-6.08	2.86-3.62	8.56-9.4
Drymaeus poecilus	31.9-38.02	7.82-8.74	3.61-4.29	23.56-28.26	6.83-7.45	3.19-3.67	10.28-12.1	2.94-3.5	0.45-0.67	16.16-20.26	14.39-17.59	16.35-20.29	10.74-13.36	4.43-4.49	1.37-1.45	6.0-6.26	3.42-4.18	7.47-8.45	9.93-11.13
Spixia tucumanensis	20.5-25.5	4.73-5.61	1.02-1.44	10.29-12.13	3.24-3.8	0.6-0.88	10.25-14.13	2.91-3.35	0.42-0.58	7.2-9.0	6.24-7.54	7.0-8.3	5.2-6.0	1.9-2.28	0.25-0.35	9.5	3.46-4.28	7.55-8.19	5.2-6.0
Plagiodontes daedaleus	23.58-28.62	5.89-6.37	2.13-2.55	15.57-19.57	4.97-5.29	1.57-1.91	9.17-11.27	2.99-3.49	0.55-0.71	11.62-14.46	11.48-14.04	10.52-12.94	8.22-10.1	3.36-3.58	0.82-0.94	6.0-7.5	3.35-4.01	9.91-10.65	6.37-7.01
Bostryx solutus	8.65-9.67	2.52-2.56	0.29-0.31	6.18-6.7	2.21-2.27	0.244-0.245	2.69-3.43	0.91-1.01	0.04-0.06	3.71-4.57	3.98-4.62	2.9-3.78	2.54-3.04	1.11-1.15	0.089-0.095	5.0	1.98-2.4	3.74-4.38	?
Bostryx conspersus	20.35-23.03	4.98-5.14	1.6-1.64	14.57-17.51	4.46-4.58	1.42-1.44	5.85-7.53	1.84-1.94	0.18-0.2	10.55-12.07	10.16-11.42	10.26-12.02	6.88-7.9	2.85-2.93	0.58-0.62	4.79-6.21	2.85-3.69	5.18-6.04	5.06-6.42
Bostryx rhodolarynx	26.41-32.37	7.07-8.11	2.73-3.99	18.4-22.77	5.77-7.01	2.17-3.41	9.0-11.02	3.04-3.2	0.55-0.57	12.9-17.66	13.48-15.03	13.48-16.24	10.43-12.13	3.9-5.12	1.13-1.95	5.79-7.21	3.83-4.73	7.3-9.4	3.56-5.12
Bostryx infundibulum	18.51-21.07	4.4-4.8	1.09-1.13	10.31-11.41	3.18-3.52	0.64-0.82	8.7-10.1	2.15-2.91	0.26-0.46	6.55-7.45	6.31-7.09	7.42-8.26	3.51-4.19	1.58-2.04	0.01-0.022	8.79-10.21	2.66-3.38	5.43-6.17	2.05-2.67
Bostryx hamiltoni	16.79-19.23	4.05-4.77	0.81-1.13	8.76-9.8	2.73-3.27	0.47-0.69	8.28-9.6	0.656-0.777	0.024-0.033	5.92-6.96	5.54-6.44	5.83-6.63	3.52-4.38	1.44-1.76	0.13-0.21	6.79-8.21	2.79-3.53	4.78-5.44	2.4-3.46
Bostryx roselleus	17.9-30.3	4.7-6.52	1.17-2.15	10.9-18.4	3.81-5.09	0.94-1.66	6.37-12.11	2.08-2.88	0.22-0.44	8.6-13.6	7.4-11.4	9.2-14.0	6.0-9.2	2.33-3.07	0.39-0.69	5.5-6.5	2.48-3.26	5.41-6.69	4.45-5.65
Bostryx birabenorum	16.08-17.4	4.330	1.191	13.84-14.72	4.0	1.126	2.89-3.93	1.291	0.079	9.28-11.38	8.27-8.83	10.23-11.41	6.1-8.48	2.693	0.501	4.25-5.01	1.33-2.25	3.1-3.96	6.42-6.8
Bostryx catamarcanus	15.77-20.19	4.3-4.5	1.1-1.25	12.23-15.97	3.84-4.14	1.02-1.19	4.23-5.51	1.32-1.39	0.09-0.1	7.75-10.49	6.76-8.76	8.29-11.79	6.11-7.73	2.61-2.95	0.49-0.62	5.5	1.2-1.9	3.7	3.04-3.46
Bostryx cordillerae	12.25-16.45	2.87-3.25	0.48-0.58	9.05-11.47	2.47-2.71	0.4-0.48	3.68-5.74	1.2-1.48	0.07-0.11	5.27-7.51	5.23-7.03	5.92-7.76	3.89-5.79	1.47-1.57	0.15-0.17	5.0-6.0	1.62-2.04	3.31-3.79	2.82-2.98
Bostryx costellatus	11.36	2.817	0.501	9.26	2.524	0.451	3.32	0.986	0.05	7.44	6.66	6.97	5.42	1.672	0.182	4.5	1.33	2.8	3.1
Bostryx cuyanus	6.21-8.85	3.17-4.49	0.65-0.99	5.38-7.04	3.04-4.44	0.56-0.92	1.59-2.95	1.21-1.55	0.05-0.11	9.42-17.68	7.82-14.56	4.56-7.36	4.61-10.31	1.47-2.09	0.11-0.17	4.0-4.5	2.73-3.03	9.93-12.63	2.36-3.5
Bostryx famatinus	17.02	3.732	0.644	10.78	2.818	0.47	6.59	1.888	0.171	5.59	5.58	7.19	4.13	1.626	0.166	7.0	2.47	3.98	3.2
Bostryx martinezi	18.3-24.6	4.95-5.41	1.29-1.55	13.8-17.7	4.04-4.6	1.06-1.34	5.4-8.8	2.06-2.16	0.2-0.24	7.89-11.01	7.27-9.71	9.7-12.7	4.98-6.98	2.21-2.97	0.31-0.55	6.0-6.26	2.36-2.92	4.73-5.35	4.44-5.4
Bostryx mendozanus	16.28-21.92	4.5-5.04	1.18-1.56	13.68-18.16	4.04-4.74	1.06-1.48	4.11-5.65	1.32-1.42	0.09-0.11	8.18-11.1	7.33-9.43	10.62-13.24	5.91-7.65	2.77-3.21	0.52-0.7	4.25-5.01	1.81-1.91	4.16-4.66	4.03-6.57
Bostryx pastorei	15.57-23.75	4.48-5.08	1.04-1.36	11.7-17.12	3.61-4.25	0.85-1.17	3.40-8.42	1.87-2.11	0.17-0.21	7.48-10.0	6.88-8.6	8.74-14.12	4.72-7.2	2.21-2.69	0.32-0.5	6.0-6.26	2.48-2.82	4.44-5.52	3.98-5.58
Bostryx peristomatus	25.2-32.5	6.43-6.97	2.06-2.8	18.1-23.2	5.37-6.27	1.76-2.52	7.6-12.6	2.14-2.38	0.26-0.3	11.7-15.8	9.4-11.8	14.5-17.6	8.4-11.8	3.66-4.76	0.94-1.62	6.0-6.5	2.71-3.57	5.39-6.17	3.34-4.4
Bostryx reedi	13.33-15.05	3.35-3.95	0.67-0.91	8.83-10.17	2.85-3.29	0.54-0.74	4.74-5.86	1.45-1.83	0.12-0.18	6.87-8.05	6.47-7.39	5.9-6.84	4.08-5.02	1.66-2.0	0.21-0.29	6.0	1.81-2.29	4.43-5.17	2.97-4.09
Bostryx scaber	25.46-31.48	7.55-8.71	3.94-5.44	22.07-26.21	7.32-8.54	3.81-5.33	5.92-7.02	2.07-2.19	0.11-0.17	19.45-23.29	15.52-18.92	17.86-21.22	13.05-14.91	5.24-6.56	1.95-3.17	5.0-6.0	2.45-3.21	6.55-7.71	6.25-7.47
Bostryx stelzneri	19.31-34.01	6.32-6.8	2.62-2.66	15.3-26.0	5.77-5.85	2.28-2.38	4.57-11.41	2.23-2.53	0.26-0.36	10.69-21.51	9.21-17.13	10.8-22.9	7.9-15.5	3.77-3.87	0.95-1.13	5.0-6.5	2.61-3.25	6.41-7.95	5.65-7.31
Bostryx strobeli	15.11-19.85	4.31-5.55	1.07-1.79	11.86-15.62	3.84-4.9	0.97-1.63	3.5-6.42	1.38-1.96	0.09-0.19	7.4-10.12	6.6-8.84	8.32-11.36	4.82-7.22	2.46	0.0889	5.0-5.5	1.61-2.79	4.35-5.61	5.55-6.15
Bostryx torallyi	25.3-32.7	5.06-6.6	1.27-2.31	15.3-22.0	3.52-5.22	0.82-1.8	8.4-14.6	2.94-3.06	0.46-0.5	9.3-13.4	8.6-12.0	10.1-15.8	5.4-8.1	1.98-3.2	0.22-0.66	7.25-8.25	3.45-4.65	7.61-9.05	6.17-7.87
Bostryx tortoranus	18.8-29.88	5.63-5.87	1.83-1.99	14.7-22.12	4.95-5.05	1.69-1.71	5.6-8.9	1.82-2.38	0.16-0.28	8.93-11.99	8.7-11.0	10.0-15.36	6.1-8.9	2.89-3.31	0.53-0.71	5.5-6.5	2.08-2.72	4.46-5.14	6.46-7.44
Bostryx willinki	17.77-19.95	4.46	0.908	11.0-12.22	3.002	0.568	7.69-9.55	2.582	0.345	6.83-8.39	6.43-7.57	7.46-8.52	4.74-5.46	1.896	0.244	6.0-7.0	2.91-3.57	5.44-5.72	3.44-3.86
Bostryx rudisculptus	12.89-16.07	2.972	0.486	9.64-11.5	2.442	0.398	4.03-5.39	1.27	0.079	5.68-6.4	5.54-5.82	7.05-7.67	4.28-4.82	1.399	0.132	5.5	1.5	3.2	3.4
Species	1	2	2	2	2	2	2	2	2	2	2	3	3	3	3	3	3	3	3	3	3	4	4	4	4	4	4	4	4	4	4
	9	0	1	2	3	4	5	6	7	8	9	0	1	2	3	4	5	6	7	8	9	0	1	2	3	4	5	6	7	8	9
Bulimulus bonariensis	0	0	0	0	0	0	0	0	0	1	0	1	0	2	0	—	0	0	—	1	0	—	0	1	1	0	0	1	1	0	2
Bulimulus apodemetes	0	0	0	0	0	0	0	0	0	1	1	1	0	1	1	0	0	0	—	1	0	—	0	1	1	0	1	1	2	0	2
Bulimulus gracilis	0	0	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	0	—	0	1	1	0	1	1	2	?	2
Naesiotus munsterii	0	0	0	0	0	0	0	0	0	1	1	1	0	0	0	—	0	0	—	2	0	—	0	1	1	0	0	1	1	0	1
Drymaeus poecilus	0	0	0	0	0	0	0	0	0	1	1	1	0	0	1	1	0	0	—	1	0	—	0	1	1	0	1	1	2	0	0
Spixia tucumanensis	0	0	0	1	0	1	1	0	0	1	0	1	0	0	1	0	1	0	—	0	1	2	1	0	0	1	—	1	0	1	0
Plagiodontes daedaleus	0	0	0	0	0	0	1	0	0	1	0	1	0	1	1	0	1	0	—	1	1	0	1	0	0	1	—	0	2	1	1
Bostryx solutus	1	0	0	0	0	0	1	0	0	1	0	0	—	—	1	0	0	1	0	?	?	?	?	?	?	?	?	?	1	0	0
Bostryx conspersus	0	0	0	0	0	0	0	0	0	1	0	0	—	—	0	—	0	0	—	1	0	—	0	?	?	?	?	?	2	0	0
Bostryx rhodolarynx	0	0	0	0	0	0	1	1	0	1	1	0	—	—	1	0	0	0	—	?	?	?	?	?	?	?	?	?	2	0	0
Bostryx infundibulum	0	1	0	0	0	0	1	0	0	1	0	0	—	—	1	0	0	1	1	?	?	?	?	?	?	?	?	?	2	0	0
Bostryx hamiltoni	0	0	0	0	0	0	0	0	0	1	0	0	—	—	1	0	0	0	—	?	1	2	?	?	?	?	?	?	2	0	0
Bostryx roselleus	0	0	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	1	1	0	1	1	—	0	1	0	1	0
Bostryx birabenorum	1	1	0	1	0	0	0	0	0	0	1	1	0	1	1	0	0	0	—	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx catamarcanus	0	0	0	0	0	0	0	0	0	1	1	1	0	1	1	0	0	0	—	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx cordillerae	0	0	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	1	1	0	1	1	—	1	1	1	0	0
Bostryx costellatus	1	1	0	0	0	0	1	0	0	0	0	1	0	1	1	0	0	0	—	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx cuyanus	0	1	0	0	2	1	1	0	1	0	0	1	0	1	1	0	0	1	0	?	?	?	?	?	?	?	?	?	1	0	0
Bostryx famatinus	0	1	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx martinezi	0	0	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	1	0	0	1	1	—	0	1	1	0	0
Bostryx mendozanus	0	0	0	0	0	0	0	0	0	0	1	1	0	1	1	0	0	0	—	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx pastorei	0	0	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	1	0	0	1	1	—	0	1	0	0	0
Bostryx peristomatus	0	1	1	0	0	0	1	1	0	1	0	1	0	1	1	0	0	1	1	1	1	2	0	1	1	—	0	1	1	1	0
Bostryx reedi	1	0	0	0	0	1	1	0	0	1	1	1	0	1	1	0	0	0	—	1	1	2	0	1	1	—	0	1	1	1	0
Bostryx scaber	1	1	0	1	1	0	0	0	1	0	1	1	1	—	0	—	0	0	—	1	1	2	0	1	1	—	0	1	1	1	0
Bostryx stelzneri	0	[01]	0	[01]	0	0	0	0	0	1	1	1	0	1	1	0	0	0	—	1	1	[01]	0	1	1	—	[01]	1	1	1	0
Bostryx strobeli	0	0	0	0	0	0	0	0	0	1	1	1	0	1	1	0	0	0	—	1	1	2	0	1	1	—	1	1	0	1	0
Bostryx torallyi	0	0	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	1	2	0	1	1	—	0	1	0	0	0
Bostryx tortoranus	0	0	0	0	0	0	0	0	0	1	1	1	0	1	1	0	0	0	—	1	1	1	0	1	1	—	0	1	1	1	0
Bostryx willinki	1	1	0	0	0	0	1	0	0	1	0	1	0	1	1	0	0	0	—	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx rudisculptus	0	1	0	0	0	0	0	0	0	1	0	1	0	1	1	0	0	0	—	1	1	0	0	1	1	—	0	1	1	0	0
Species	5	5	5	5	5	5	5	5	5	5	6	6	6	6	6	6	6	6	6	6	7	7	7	7	7	7	7	7
	0	1	2	3	4	5	6	7	8	9	0	1	2	3	4	5	6	7	8	9	0	1	2	3	4	5	6	7
Bulimulus bonariensis	0	1	0	0	0	1	1	1	1	0	0	0	0	—	0	0	1	0	0	0	0	0	1	1	0	1	0	1
Bulimulus apodemetes	0	1	1	0	0	1	1	1	0	0	0	0	0	—	0	0	1	0	0	0	0	1	[01]	1	0	0	0	1
Bulimulus gracilis	1	0	1	0	0	1	1	1	1	0	0	0	0	—	0	0	1	0	?	0	0	0	1	1	0	0	0	1
Naesiotus munsterii	1	0	0	0	0	1	1	2	0	0	0	0	0	—	0	0	1	0	0	0	0	0	1	1	0	0	0	1
Drymaeus poecilus	0	1	0	0	?	0	1	2	0	0	0	0	0	—	0	0	1	0	1	0	0	0	2	1	0	1	0	1
Spixia tucumanensis	1	0	0	0	1	0	1	0	0	0	0	2	0	—	0	1	1	1	0	0	1	0	2	1	1	1	1	0
Plagiodontes daedaleus	1	0	0	0	0	1	1	0	0	0	1	0	0	—	0	0	1	1	1	0	1	0	2	0	1	0	1	0
Bostryx solutus	?	?	0	?	1	?	1	2	?	?	?	?	?	?	?	?	?	?	?	?	0	?	?	1	0	2	0	1
Bostryx conspersus	?	?	1	0	1	1	0	—	—	?	?	?	?	?	?	?	?	—	?	?	0	0	1	1	0	1	0	1
Bostryx rhodolarynx	?	?	1	0	1	?	1	2	?	?	?	?	?	?	?	?	?	?	?	?	0	1	0	1	0	1	0	1
Bostryx infundibulum	?	?	0	0	1	?	1	2	?	?	?	?	?	?	?	?	?	0	?	?	0	0	2	1	0	1	0	1
Bostryx hamiltoni	?	?	0	0	1	?	1	2	?	?	?	?	?	?	?	?	?	0	?	?	0	?	?	1	0	1	0	1
Bostryx roselleus	0	1	0	1	1	1	1	2	1	1	0	1	1	2	1	0	0	0	1	1	0	1	0	1	0	2	0	1
Bostryx birabenorum	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx catamarcanus	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx cordillerae	0	1	0	1	1	1	1	2	1	1	0	1	1	0	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx costellatus	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx cuyanus	?	?	0	1	1	?	1	2	?	?	?	?	?	?	?	?	?	0	?	?	0	1	0	1	0	2	0	1
Bostryx famatinus	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx martinezi	0	1	0	1	1	1	1	2	1	0	0	1	1	0	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx mendozanus	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx pastorei	0	1	0	1	1	1	1	2	1	0	0	1	1	0	1	0	0	0	1	1	0	1	0	1	0	2	0	1
Bostryx peristomatus	0	1	0	1	1	1	1	2	1	0	0	1	1	0	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx reedi	0	1	1	1	1	1	1	2	1	0	0	1	1	1	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx scaber	0	1	1	1	1	1	1	2	1	0	0	1	1	[01]	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx stelzneri	0	1	1	1	1	1	1	2	1	1	0	1	1	[01]	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx strobeli	0	1	1	1	1	1	1	2	1	0	0	1	1	0	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx torallyi	0	1	0	1	1	1	1	2	1	0	0	1	1	1	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx tortoranus	0	1	1	1	1	1	1	2	1	0	0	1	1	1	1	0	0	0	0	1	0	1	0	1	0	2	0	1
Bostryx willinki	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?	?
Bostryx rudisculptus	0	1	1	1	1	1	1	2	1	0	0	1	1	0	1	0	0	0	1	1	0	1	0	1	0	2	0	1

Museu de Zoologia da Universidade de São Paulo Av. Nazaré, 481, Ipiranga, 04263-000 São Paulo SP Brasil, Tel.: (55 11) 2065-8133 - São Paulo - SP - Brazil
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