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(Updated: 2022/01/31)

About the journal


Basic Information


Hoehnea is currently the scientific journal of the Instituto de Pesquisas Ambientais, published by the Institute of Botany since 1971, It superceded Archivos de Botânica de São Paulo. Hoehnea publishes continuously, from 2020, original articles, reviews and scientific notes on all aspects of plant and fungal sciences, in portuguese, spanish and mainly in english. The journal name honors botanist Frederico Carlos Hoehne, founder and first director of Instituto de Botânica. The journal is available online on open access, since volume 29 at:

Access Policy - Hoehnea is published under the Open Access model. There are not fees for the processing of articles.

Hoehnea Journal does not charge any type of fees from the authors.

Its abbreviated title is Hoehnea, which should be used in bibliographies, footnotes and bibliographical references and strips.



Indexed in



Intellectual Property


All content of the journal, except where identified, is licensed under a Creative Commons attribution-type BY.





The publication receives funding from


Instituto de Pesquisas Ambientais

Av. Miguel Stefano, 3687, Água Funda, São Paulo/SP, Brazil, 04301-902




Editorial Board




Maria Margarida da Rocha Fiuza de Melo (Instituto de Botânica, São Paulo, SP, Brasil)



Assistant Editors


Adriana de Mello Gugliotta (Instituto de Botânica, São Paulo, SP, Brasil)

Claudio José Barbedo (Instituto de Botânica, São Paulo, SP, Brasil)

Eduardo Pereira Cabral Gomes (Instituto de Botânica, São Paulo, SP, Brasil)

Iracema Helena Schoenlein-Crusius (Instituto de Botânica, São Paulo, SP, Brasil)

Regina Maria de Moraes (Instituto de Botânica, São Paulo, SP, Brasil)



Associated Editors


Abel Sentíes (Ficologia: algas marinhas - Universidad Autónoma Metropolitana, campus Iztapalapa, Ciudad de México, México)

Adriana Hissae Hayashi (Anatomia Vegetal - Instituto de Botânica, São Paulo, SP, Brasil)

Alain Chautems (Taxonomia de Plantas Vasculares - Conservatoire et Jardin Botaniques de la Ville de Genève, Genebra, Suiça)

Alice da Cruz Lima Gerlach (Liquenologia e Micologia – Genève Botanical Garden, Pregny-Chambésy, Suíça)

Ana Paula de Souza Caetano (Anatomia Vegetal – Universidade Federal de Mato Grosso, Cuiabá, MT, Brasil)

Ana Rita Giraldes Simões (Taxonomia de Plantas Vasculares – Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AE, United Kingdom)

André Aptroot (Liquenologia e Micologia – Universidade Federal do Mato Grosso do Sul, Pioneiros, MS, Brasil)

Anna Luiza Ilkiu Borges Benkendorff (Briologia e Pteridologia – Museu Paraense Emílio Goeldi, Belém, PA, Brasil)

Bárbara Dunck Oliveira (Ficologia: algas de água doce – Universidade Federal Rural da Amazônia, Belém, PA, Brasil)

Carlos Eduardo Wetzel (Ficologia: algas de água doce – Luxembourg Institute of Science and Technology, Esch-sur-Alzette, Luxemburgo)

Catarina Carvalho Nievola (Fisiologia e Bioquímica Vegetais – Instituto de Botânica, São Paulo, SP, Brasil)

Cláudia Baider (Ecologia - The Mauritius Herbarium, Réduit, Ilha Mauricio)

Claudia Elena Carneiro (Ensino em Botânica e Educação Ambiental – Universidade Estadual de Feira de Santana, Feira de Santana, BA, Brasil)

Cynthia Fernandes Pinto da Luz (Palinologia – Instituto de Botânica, São Paulo, SP, Brasil)

Diego Tavares Vasques (Briologia e Pteridologia – The University of Tokyo, Tokyo, Japan)

Enrique Forero (Taxonomia de Plantas Vasculares - Academia Colombiana de Ciencias Exactas Fisicas y Naturales, Bogotá, Colombia)

Fátima Otavina de Souza Buturi (Taxonomia de Plantas Vasculares - Universidade São Judas Tadeu, São Paulo, SP, Brasil)

Francisco de Assis Ribeiro dos Santos (Palinologia - Universidade Estadual de Feira de Santana, Feira de Santana, BA, Brasil)

Giselda Durigan (Restauração Ecológica - Instituto Florestal do Estado de São Paulo, Assis, SP, Brasil)

Gisele Carolina Marquardt (Ficologia: algas de água doce – Instituto de Botânica, São Paulo, SP, Brasil)

Juliana Hanna Leite El Ottra (Anatomia Vegetal – Universidade de São Paulo, São Paulo, SP, Brasil)

Natália Macedo Ivanauskas (Ecologia - Instituto Florestal, São Paulo, SP, Brasil)

Nelson Augusto dos Santos Júnior (Ecofisiologia de sementes – Instituto de Botânica, São Paulo, SP, Brasil)

Paulo Silveira (Taxonomia de Plantas Vasculares – Universidade de Aveiro, Aveiro, Portugal)

Rafael Batista Louzada (Taxonomia de Plantas Vasculares – Universidade Federal de Pernambuco, Recife, PE, Brasil)

Regina Hirai (Briologia, Pteridologia - Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rio de Janeiro, RJ, Brasil)

Renata Sebastiani (Taxonomia de Plantas Vasculares – Universidade Federal de São Carlos, Araras, SP, Brasil)

Renato Crespo Pereira (Ficologia: algas marinhas - Universidade Federal Fluminense, Niterói, RJ, Brasil)

Rosana Maziero (Liquenologia e Micologia - Comitato scientifico del Gruppo Micologico Milanese, Milano, Italia

Viviana Motato-Vásquez (Liquenologia e Micologia – Universidad de Valle, Cali, Colombia)

Viviane Stern da Fonseca Kruel (Etnobotânica – Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rio de Janeiro, RJ, Brasil)

Wagner Melo Ferreira (Fisiologia Vegetal - Universidade Federal do Tocantins, Porto Nacional, TO, Brasil)



Editorial Production


Karina Margaret Silva das Neves (Editoração Técnica) –  Instituto de Botânica, São Paulo, SP, Brasil

Maria Margarida da Rocha Fiuza de Melo (Editoração Técnica) – Instituto de Botânica, São Paulo, SP, Brasil



Instructions to authors


Scope and Policy


Hoehnea publishes original complete articles, reviews, and scientific notes in all fields of Botany and Mycology (anatomy, cell and molecular biology, biochemistry, ecology, phylogeny, physiology, genetics, morphology, palynology, taxonomy) in Portuguese, Spanish, or English. The review articles are exceptionally accepted at the discretion of the Editorial Board and must contain a critical analysis of the main subject and shouldn't be restricted to bibliographic compilations. Scientific notes must present relevant technical and scientific approaches.

The author has to submit your manuscript in the ScholarOne System, containing the identification of the authors, title, and full address of the all authors. The corresponding author have to suggest possible reviewers (with their contacts) or request that others may be excluded. The authors should express their agreement to transfer the copyright to the Instituto de Pesquisas Ambientais and declare that the manuscript was not published, in whole or in part, and not submitted to another journal. The submission of the manuscript implies the author's commitment that it was not and will not be published elsewhere, in paper or electronic format, in domestic or foreign publications.

Manuscripts submitted for review should consist of a single document in Word (.doc, .docx, or .rtf) with tables and figures with low resolution (150 dpi) attached at the end of the document. The submission can be done by Sistema de Submissão Eletrônica ScholarOne ( Accepted for publication, the Editor-in-Chief will request high-resolution tables and illustrations (600 dpi) separately.

Open Access Policy - Hoehnea is published under the Open Access model and is therefore free for anybody to read and download, and to copy and disseminate for educational and scientific purposes.



Form and preparation of manuscripts


General Information - The original manuscripts should be sent to the editor-in-chief and should be in accordance with the Instructions to Authors. Manuscripts that do not meet the specifications set in the Instructions are immediately returned to the author(s) for revisions.
The manuscripts that are in accordance with the Instructions to Authors will be sent to the associate editors indicated by the editor-in-chief. In any case, the evaluation will be anonymously sent to the authors. The manuscripts will be published according to the chronological order of acceptance by the Editorial Board and not by the receiving order.

Manuscript format - use Word for Windows 6.0 or a latest version, Times New Roman font, size 12, double-spaced. Text aligned for the right margin and not justified. Use A4 paper (297 × 210 mm) with 2 cm for all margins. Number pages consecutively and avoid footnotes. Manuscript should not exceed 50 typed pages, including tables and figures. Scientific notes should not exceed ten pages.

Insert line numbers to facilitate review comments by Reviewers, Associate Editors, and the Editor-in-Chief.

First page - title in bold and only the initials in capital letters; author's full name (at author's discretion), capitalizing initials and using lowercase for the other letters; affiliation, complete addresses, and corresponding author's e-mail should be as footnotes, indicated by numbers; and the shortened title and ORCIDs numbers of all authors. If the manuscript is part of a Thesis, Dissertation, or Work for completion of course, indicate it by using a superscript numeral in the title. Grants, scholarship, and number of process from financial agency should be included in the Acknowledgements. After the manuscript goes through the scientific publishing process, changes in the list of authors are not accepted. The contacts with the Editorial Board of Hoehnea can be made only through the corresponding Author.

The manuscript should contain only the essential information for its understanding, and be rigorously prepared according to the guidelines.

Second page - should contain ABSTRACT and RESUMO (or RESUMEN), beginning with the title of the paper in the corresponding language between parentheses. It should not exceed 150 words in a single paragraph. Keywords and Palavras-chave (or Palabras clave) should be up to five words separated by comma, without period, and in alphabetic order. Do not use words that have been used in the title.

Text - start in a new page. Chapter titles should be written in bold, centralized, and only the initials in capital letters, following as possible: Introduction, Material and methods, Results, Discussion, Acknowledgments, and Literature cited. Results and Discussion may be combined in one. Subtitles of these topics should be written with regular font with uppercase and lowercase letters and separated from the corresponding text with a hyphen. Use italic for scientific names. After the Acknowledgments item, if any, include two items: Conflict of interests and Authors' contributions - contributions in the preparation of the manuscript being submitted.

Abbreviations of books and the taxon's author or authors - should be according to Brummitt & Powell (1992). For taxonomic papers, in the headers of the species, abbreviations of rare books must follow the Taxonomic Literature (TL-2) and for periodicals, the Botanico-Periodicum-untianum/Supplementum (B-P-H/S).

Figures and tables - must be consecutively numbered with Arabic numerals following the order in which they are cited in the text. Do not use abbreviations when referring to them in the text. For taxonomy papers, figures should be cited as follow:

Bauhinia platypetala Burch. ex Benth. in Mart, Fl. Bras. 15(2): 198. 1870 = Bauhinia forficata Link var. platypetala (Burch. ex Benth.)
Wunderlein, Ann. Missouri Bot. Gard. 60(2): 571. 1973. Tipo: BRASIL. Tocantins: Natividade, s.d., G. Gardner 3118 (síntipo OFX).
Figuras 7-8

References in the text - should be cited using the author-date System, with initials in capital letters; when in the same set of citations, use chronological order; for two authors, link names with &; for more than two, mention the first, followed by et al.; for papers published in the same year by the same author or the same combination of authors, use letters following the publication year (e.g.: 1994a, b); do not use comma to separate author and publication year, rather, use it to separate multiple citations (ex.: Dyer & Lindsay 1996, Hamilton 1988); references to unpublished results should be as follows: (Capelari, unpublished data).

Herbarium specimens - herbarium material must be cited according to the following model: BRASIL. São Paulo (written with versalete effect – small caps): São Paulo, São Paulo, Parque Estadual das Fontes do Ipiranga, collection date (e.g.: 10-IX-1900), collector name plus collection number (herbarium acronym). Cite only the herbarium acronym when a collector number is cited; when collector number is not provided, cite the herbarium acronym followed by the herbarium registration number [(e.g.: (SP250874)]. Use the abbreviation s.d. when the date is not provided.

Numbers and units - use abbreviations of measurements units whenever possible; for composed units use exponentials instead of bars (ex.: mg day-1 instead of mg/day, µg L-1 instead of µg/L), leaving a space between number and unit (ex. 200 g; 50 m); do not insert spaces in coordinates (ex.: 23o46'S and 46o18'W).

For approximate measure use circa (ca. 5 cm); for a range of measures do not use circa (2-5 cm); for approximate form use ca. (ca. 3 × 5 cm); for temperature (20 oC); for a percentage value only (60%); variation range to percentages (30 - 50%).

Literature cited - list of the authors in bold letters and in alphabetical order, capitalizing only the initials; citations of the same author or combination of authors should follow chronological order for a set of references with the same first author; work of the author alone must be cited first then the author with one co-author, and finally the author with two or more co-authors. Full periodical names should be used. Avoid citing dissertations and thesis; do not cite conference abstracts, graduation monograph, or articles in press. Authors should follow the examples:

Manuscripts of Meetings
Giannotti, E. & Leitão Filho, H.F. 1992. Composição florística do cerrado da Estação Experimental de Itirapina (SP). In: R.R. Sharif (ed.). Anais do 8º Congresso da Sociedade Botânica de São Paulo, Campinas, pp. 21-25.

Articles of periodicals
Pôrto, K.C., Gradstein, S.R., Yano, O., Germano, S.R. & Costa, D.P. 1999. New an interesting records of Brazilian bryophytes. Tropical Bryology 17: 39-45.
Veasey, E.A. & Martins, P.S. 1991. Variability in seed dormancy and germination potential in DesmidiumDesv. (Leguminosae). Revista de Genética 14: 527-545.

Cronquist, A. 1981. An integrated system of classification of flowering plants. 2 ed. New York Botanical Garden, New York.
IPT. 1992. Unidades de conservação e áreas correlatas no Estado de São Paulo. Instituto de Pesquisas Tecnológicas, São Paulo.
Chapters in books and serial works
Benjamin, L. 1847. Utriculariae. In: C.F.P. Martius (ed.). Flora Brasiliensis. Typographia Regia, Monachii, v. 10, pp. 229‑256, t. 20‑22.
Ettl, H. 1983. Chlorophyta, I. Phytomonadina. In: H. Ettl, J. Gerloff, H. Heynig& D. Mollenhauer (eds.). Süswasser Flora von Mitteleuropa, Band 9. Gustav Fischer Verlag, Sttutgart, pp. 1-809.
Heywood, V.H. 1971. The Leguminosae - a systematic review. In: J.B. Harbone, D. Boulter & B.L. Turner (eds.). ChemotaxonomyoftheLeguminosae. Academic Press, London, pp. 1-29.

Electronic documents
Poorter, H. 2002. Plant growth and carbon economy. Encyclopedia of Life Sciences. Available at (access in 20-XI-2004).

Thesis or Dissertations
Trufem, S.F.B. 1988. Fungos micorrízicos vesículo-arbusculares da Ilha do Cardoso, SP, Brasil. Tese de Doutorado, Universidade de São Paulo, São Paulo.

Books with coordinator, editor or organizer
Mamede, M.C.H., Souza, V.C., Prado, J., Barros, F., Wanderley, M.G.L. & Rando, J.G. (orgs.). 2007. Livro vermelho das espécies vegetais ameaçadas do Estado de São Paulo. Instituto de Botânica, São Paulo.


Secretaria do Meio Ambiente do Estado de São Paulo. 2004. Resolução SMA-48, de 21-IX-2004. Lista de espécies da flora ameaçadas de extinção no estado de São Paulo. Diário Oficial do Estado de São Paulo, Poder Executivo, São Paulo, 22-IX-2004. Seção I, v.114, n. 179, pp. 26-29.

Works with corporate authors
ABNT. 2002. NBR 6023: Informação e documentação – Referências – Elaboração. Associação Brasileira de Normas Técnicas, Rio de Janeiro.

Tables - use design resources and table formatting of Word for Windows and present tables in separate pages. Do not insert vertical lines; horizontal lines should be used only in headings and to close the table. Titles should be concise and inserted in the upper part, starting with “Table”, followed by Arabic numbers in the order in which they are mentioned in the text. Avoid abbreviations (except for units) but if not possible, add its meaning in the title. For tables covering more than one page, add “continue” at end of the Table and “Table 1 (continuation)” in the upper left corner of the following page, repeating the heading but not the caption. Manuscripts in Portuguese or Spanish should also include table captions in English.

Figures - Figures or group of figures should be presented in separate archives; titles should be listed in sequence at the end of the manuscript. Each figure (photograph, drawing, graph, map, or scheme) should be numbered with Arabic numerals in the order of which they are mentioned in the text; small letters can be used to subdivide figures (dark figure use letters with light color and the opposite in light figures); figure numbers or letters should be placed, whenever possible, on the low right corner (use Times New Roman font). In the bar graphs indicate the conventions of the bars in the figure title. The maximum length of each figure or group of figures should be 230 mm, including the title; width can be adjusted to one or two columns (81 or 172 mm). The originals should be prepared on the final size or be proportional to a maximum of two times the publication size (the dimensions of the printed page are 230 mm long and 172 mm wide). Plates containing original ink drawings should be presented on a good quality white/drawing paper. Lines and letters must be sharp enough to allow reduction. Only good quality, black and white figures will be accepted. Magnifications (metric system) must be indicated with scale bars clearly shown in the left side of the figures. Use Times New Roman font in the titles of figures and graphics. Digital figures are accepted in .tif format with a minimum resolution of 600 dpi and must not be embedded in MS Word. Figures with low quality or out of proportion will not be accepted. Manuscripts in Portuguese or Spanish should also include figure title in English.

Additional information

In case of doubts about the instructions, it is recommended that authors consult a recent article published in Hoehnea, in the same area of knowledge of the manuscript. All articles are reviewed by at least two referees in the area of knowledge.

The Editor-in-Chief, assistant editors, with the assistance of the referees, will decide about the acceptance for publication, and will advise modifications in the articles.

For articles accepted with modification, the authors should respond to the reviewers' comments, one by one, in a table with two columns: the left with the review of the evaluator and the right with the reply from the author, justifying the acceptance or non-acceptance of the suggestions.

The authors are responsible for the opinions and concepts emitted in the article.

These instructions are also available at the website of the Instituto de Pesquisas Ambientais ( and at SciELO - Scientific Electronic Library Online (

Additional information

1. Hoehnea Journal publishes in the continuous flow mode each volume of the year).

2. Hoehnea Journal can change the format and make grammar corrections in the manuscript to accomodate it to Hoehnea's editorial policy and linguistic approach. The final proof is sent to the authors for a final review. At this stage, only the typographical and orthographic erros can be corrected.

3. Hoehnea Journal does not charge any type of fees from the authors. Additional information can be obtained by e-mail at For information about the status of a manuscript, provide the submission number.

4. Plagiarism Policy: the Hoehnea Journal doesn't accept plagiarism anyway. We rely on the help of the Editorial Board to investigate possible serious bad behavior. And informed or detected plagiarism, at any stage, will be investigated and the manuscript will be removed immediately, with vehement rebuke to the authors.

5. Copyright: when forwarding a manuscript, authors should be aware that if accepted for publication, the article's copyright should be given exclusively to the Journal Hoehnea.



Send of the manuscripts


Manuscripts should consist of a single document (doc or rtf), tables with figures in low resolution (150 dpi) attached at the end of the document.

After review, once accepted for publication, the editor will request tables and original artwork in high resolution. The maximum height to figure or figures on the printing group is 230 mm, including the label, can be adjusted to the width of one or two columns (81 mm or 172 mm), in proportion (up to twice) to the final area of the occupation of the figure (the dimensions of the magazine is 230 mm high by 172 mm wide). The drawings must be original, made with black ink on white paper of good quality or plant, lines and letters must be clear, allowing reduction. Photos and graphics are accepted in black and white, and colored. The scale used is the metric and should be plotted on the left side of the figure. Use Times New Roman in the legends of figures and graphs. Accepted figures digitized, providing they have clarity and be sent in. Tif with at least 600 dpi graphics, and should not be embedded in MS Word or Power Point.




Instituto de Pesquisas Ambientais Av. Miguel Stefano, 3687 , 04301-902 São Paulo – SP / Brasil, Tel.: 55 11 5067-6057, Fax; 55 11 5073-3678 - São Paulo - SP - Brazil