ISSN 0031-1049 printed version
ISSN 1807-0205 online version



General Information

Papéis Avulsos de Zoologia (PAZ) covers primarily the areas of Zoology, publishing original contributions in systematics, paleontology, evolutionary biology, ecology, taxonomy, anatomy, behavior, functional morphology, molecular biology, ontogeny, faunistic studies, and biogeography.

The journal also encourages the submission of theoretical and empirical studies that explore the principles and methods of systematics.

All contributions must follow the rules and recommendations of the International Code of Zoological Nomenclature (ICZN).

Relevant specimens must be deposited in national or international public museums or collections.

Tissue samples should be referred to their voucher specimens and all nucleotide sequence data (aligned as well as unaligned) should be submitted to GenBank or European Molecular Biology Laboratory (EMBL).

ZooBank Registration:

ZooBank is the Official Register of the International Commission on Zoological Nomenclature (ICZN) for registration of new nomenclatural acts, published works, and authors.
The editorial team of Papéis Avulsos de Zoologia is going to register articles that contain nomenclatural acts in ZooBank. Such articles will receive a Life Science Identifier (LSID) that is going to be included in the published version.

The registration of new names in Zoobank is recommended (not a requirement) to authors.


Peer Review

All submissions are subject to analysis by at least 2 (two) referees qualified to evaluate the manuscript and one Editor.

The peer review type adopted is “single-blind”, in which the names of the reviewers are hidden from the author.

Once a manuscript is accepted providing changes suggested by the referees, the author is requested to return a revised version incorporating those changes (or a detailed explanation of why reviewer's suggestions were not followed) within a maximum of 15 (fifteen) days upon receiving the communication by the Editor.



Page-proofs with the revised version will be sent by email to the corresponding author.

Page-proofs must be returned to the Editor, preferentially within 48 (forty-eight) hours.

Only necessary corrections in proof will be permitted.

The omission of return the proof will mean automatic approval of version with no changes and may result in a delay in publication.

Article Processing Charge (APC)

There is no publication fee to submit or publish in Papéis Avulsos de Zoologia.


Submission of Manuscripts

IMPORTANT: You must correctly complete all fields for each AUTHOR: Name, Middle Name, Last Name, Email, ORCID, URL (if there is), Institution/Affiliation, Country, and Biography Summary.

From 2019 on, the inclusion of the ORCID ID for each author is mandatory.

The fields for the manuscript should also be filled with the information: Title, Abstract, Area and Sub-Area of Knowledge, Keywords, Language and Funding/Support Agencies.

  • The manuscript must be sent only in ENGLISH.
  • Manuscripts should be sent in the format “.DOCX” or “.RTF” to the journal submission system, along with a submission letter explaining the importance and originality of the study.
  • All submissions are performed through the “Portal de Revistas da USP” .
  • Pictures, graphics, and illustrations even inserted in the text must necessarily be sent in separate files, in the original format in which they were created.
  • The files must be sent in separate files formats: “.PSD”, “.TIF”, “.JPG” “.PCX”, “.GIF” or “.BMP” for figures; “.EPS”, “.CDR”, “.WMF”, “.AI”, “.PPTX” or “.XLSX” for graphics and “.MOV”, “.AVI”, “.MPG”, “.MP4” or “.M4V” for videos.
  • Tables should be placed at the end of the manuscript.
  • Always keep updated the email and the address of the author(s) or corresponding.
  • The authors may suggest potential reviewers.



Manuscript Form

• The manuscripts must be double-spaced between lines, justified text, and Calibri font, body 11 (eleven) (except for special symbols not included in this font).

• The text should be organized in the following sections, each started on a new page, in the order, and numbered consecutively: Title Page, Abstract with Keywords, Body Text, Acknowledgments, References, Appendices, Tables, and Figure Legends.

•  Scientific names of species and genera and other Latin terms must be italicized in all sections of the manuscript.

(1) Title Page: Should include the Title, Running Title, Author(s) Name(s), ORCID numbers(s), Institution(s) and Address(es). The title should be concise and, where appropriate, should include particulars about families and/or taxa of higher categories. New taxa names should not be included in the titles.

(2) Abstracts: All papers should have an abstract and keywords in English. The quality of the abstract is of great importance since it can be reproduced in other vehicles. Therefore, must be written in an intelligible form as it may be published separately, and should summarize the main facts, ideas, and conclusions of the article. Telegraphic abstracts are unacceptable. Finally, the abstract should include all new taxonomic names for reference purposes. Abbreviations should be avoided. It should not include bibliographical references. Abstracts and keywords must not exceed 350 (three hundred and fifty) and 5 (five) words, respectively.

(3) Body Text: The main body text varies with different types of papers, but should usually include the following sections: Introduction, Material and Methods, Results, Discussion, Conclusion, Acknowledgements and References. The main headings of each section of the body of the text should be written in capital letters, bold and centered. Secondary headings should have only the first letter capitalized, bolded, and centered. Tertiary headings should be the first letter capitalized, in bold and indented left. In all cases, the text should start on the next line.

(4) References: Citations in the text should be given as follows: Martins (1959) or (Martins, 1959) or (Martins, 1959, 1968, 2015) or Martins (1959a, b) orMartins (1959: 14-20) or Martins (1959: figs. 1, 2) or Martins & Reichardt (1964) or (Lane, 1940; Martins & Chemsak, 1966a, b) or Martins et al. (1966) or(Martins et al., 1966), the latter when the article contains 3 (three) or more authors. There is no need to provide a full reference when authors and date are presented only as an authority of taxa.

(5) References Models: References should be arranged alphabetically and according to the following format, respecting the titles in italics:

  • Journal Article - Author(s). Year. Article title. Journal name, volume: initial page-final page. Journal titles must be written in full (not abbreviated).
  • Books - Author(s). Year. Book title. Publisher, Place of publication.
  • Books Chapters - Author(s). Year. Chapter title or part. In: Author(s) or Editor(s), Book title. Publisher, Place of publication, volume or part, initial page-final page.
  • Dissertations and Theses - Author(s). Year. Title of dissertation or thesis. (Masters Dissertation or Doctoral Thesis). University, Place of publication.
  • Electronic Publications - Author(s). Year. Document title. Available at: electronic address. Accessed : DD/MM/AAAA.
  • Datasets - Author(s). Year. Title. Version. Publisher. [dataset]. Available at: (preferably the DOI number as url). Accessed: DD/MM/AAAA. - See more in the Research Data section.


  • All tables should be numbered in the same sequence in which they are cited in the text.
  • Legends should be self-explanatory, without the need to resort to text.
  • Tables should be formatted preferentially towards portrait, remaining the sense landscape for exceptional cases.
  • In the text, tables should be referred as Table 1, Tables 2 and 4, Tables 2-6.
  • Use “TABLE” in the table(s) heading(s).


  • The figures must be numbered consecutively, in the same sequence in which they are cited in the text.
  • Each illustration of a composite figure should be identified by a capital letter and referred to in the text as, for example: Fig. 1A, Fig. 1B.
  • When possible, the letters must be positioned in the lower-left corner of each illustration of a composite figure.
  • Photographs in black and white or color must be scanned at high resolution (300 DPI minimum).
  • Use “Fig(s).” for referring to figures in the text, “FIGURE(S)” in the figure captions and “fig(s).” when referring to figures in another paper.
  • The figures will be printed in black and white but maintained in color in the final PDF file.

Third-party content:
Previously published content as figures, tables, illustrations, etc. must be accompanied by written permission from the copyright holder to reproduce.

Responsibility: The scientific content and opinions expressed in this publication are sole responsibility of the respective authors.

Authorship and Authors' Contributions Statement:
All Authors must meet the two minimum criteria:

1) actively participate in the results discussion;
2) review and approve the final version of the paper.

An authors' contributions statement must be included in a section at the end of the article for works written by more than 4 (four) authors. The statement has to include the individual contribution of each author.
Acknowledgments section may contain a declaration of other kinds of contributions.

Copyrights: A Term of Assent and Cession of Copyrigh (attached template) should be signed and sent to the Editor, prior to publication of the manuscript.
Papéis Avulsos de Zoologia adopts for publication the Creative Commons License – CC-BY (

Conflict of Interest:
The author must declare if there is any conflict of interest at the submission of the manuscript.
If there is no conflict of interest, the authors should select in the submission form the option: "Authors declare that there is no conflict of interest."; otherwise, it must be specified in "Comments to the Editor".
Coeditors, Associated Editors, and Reviewers must declare to the Editor any conflict of interest before starting the review process.
For further information on Conflict of Interest refer to the Council of Science Editors White Paper on Publication Ethics (English or Portuguese versions).

Funding information:
Research grants from funding agencies (please inform the research funder and the grant number) or any financial support must be declared.


For more details of the manuscript preparation format, see CBE Style Manual, available at the electronic address of the Council of Science Editors (

Papéis Avulsos de Zoologia is a publication of the Museu de Zoologia da Universidade de São Paulo (

Always consult the Instructions to Authors printed in the last issue or in the electronic home pages: or


Research data

Papéis Avulsos de Zoologia recommends the deposit of scientific data in appropriate repositories and its citation in accordance with FAIR principles (Findable, Accessible, Interoperable and Re-usable).

As described in the “General Information” section of this Instructions to Authors, all nucleotide sequence data (aligned as well as unaligned) should be submitted to  GenBank or European Molecular Biology Laboratory (EMBL). All articles that contain nomenclatural acts are registered in Zoobank by the journal staff and the Life Science Identifier (LSID) of the article is included in the published version.

Authors are encouraged to deposit their data in a repository that is best suited to their dataset. Some examples of repositories are the Global Biodiversity Information Facility (GBIF), Environmental Data Initiative (EDI), Dryad, Figshare, Zenodo, etc.  

The data repository used must guarantee the preservation of the data and provide a persistent identifier such as a DOI so it can be accessible and citable.

Journal will make exceptions for sensitive information as the location of endangered species.

Datasets used in the research, deposited in a scientific research data repository, should be cited in the “Materials and method section” and its reference (with the DOI number) should be included in the reference list.

Datasets References Examples:

  • Dataset deposited in a scientific research data repository:

Botham, M.; Roy, D.; Brereton, T.; Middlebrook, I.; Randle, Z. 2013. United Kingdom Butterfly Monitoring Scheme: species trends 2012. NERC Environmental Information Data Centre. [dataset]. Available at: . Accessed: 13/04/2019.

United States Department of Health and Human Services. Substance Abuse and Mental Health Services Administration. Office of Applied Studies. 2015. Treatment Episode Data Set -- Discharges (TEDS-D) -- Concatenated, 2006 to 2011. Version V5. Ann Arbor, MI: Inter-university Consortium for Political and Social Research [distributor], 23 Nov. 2015. Available at: . Accessed: 30/09/2019.

  • Article supplementary information deposited in a scientific research data repository:

Yoon, J; Sofaer, H.R, Sillet, T. S, Morrison, S.A., Ghalambor, C.K. 2017. Data from: The relationship between female brooding and male nestling provisioning: does climate underlie geographic variation in sex roles?. Journal of Avian Biology, 48(2):220-228. Available at: . Accessed: 07/07/2019.


  • Dataset published as data paper:

Bovendorp, R. S., Villar, N., de Abreu-Junior, E. F., Bello, C., Regolin, A. L., Percequillo, A. R., & Galetti, M. 2017. Atlantic small-mammal: a dataset of communities of rodents and marsupials of the Atlantic forests of South America. Ecology, 98(8):2226–2226. [data paper]. Available at: Accessed: 07/10/2019.


Author Self-Archiving

Authors can share the published version* of the manuscript with their colleagues and post them on personal websites or institutional repositories for academic purposes while providing bibliographic details that credit, if applicable, its publication in this journal.

*Published version - is the article published in PDF format available at the Papéis Avulsos de Zoologia website.


Publication Ethics

Research or publication misconduct (plagiarism, self-plagiarism, fabrication or falsification of data and results, etc.) will be treated in accordance with the Commission on Publication Ethics (COPE) guidelines.

From May 2020 on, the journal has adopted the tool Originality Check (Turnitin) to prevent duplication and plagiarism occurrences.

Allegations, corrections, and doubts must be sent to the Editor (contact information). All allegations/cases will be evaluated and, when necessary, retractions, corrections or expressions of concern will be published.


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