Open-access An illustrated catalog of Disteniidae (Coleoptera: Chrysomeloidea) type specimens deposited at the Museu de Zoologia da Universidade de São Paulo

Abstract

The type specimens of the family Disteniidae Thomson, 1861 (Coleoptera) housed at the Museu de Zoologia, Universidade de São Paulo (MZSP) are cataloged and illustrated. Photos of dorsal and ventral habitus, labels, and complete label data of all the specimens are provided. There are 147 specimens belonging to 70 names: 33 primary types, 31 holotypes, and two lectotypes; and 114 secondary types, 52 paratypes, and two paralectotypes. Elytrimitatrix (Grossifemora) Santos-Silva & Hovore, 2007 is the genus with the greatest number of species with type material, 21 species, followed by Cometes Lepeletier & Audinet-Serville, 1828 and Novantinoe Santos-Silva & Hovore, 2007 with 15 and 13 species, respectively.

Keywords
America; Cerambycoidea; Entomological collection; Neotropical Region

INTRODUCTION

The family Disteniidae is currently composed of more than 410 species (Monné, 2023; Tavakilian & Chevillotte, 2023) and is distributed globally, predominantly in tropical and subtropical regions (Švácha & Lawrence, 2014). According to Švácha & Lawrence (2014), the family is divided into four tribes: Cyrtonopini, Disteniini, Dynamostini, and Heteropalpini.

Disteniidae, together with Cerambycidae, Oxypeltidae, and Vesperidae, are grouped into the so-called “Cerambycoidea” clade, a group which is part of the superfamily Chrysomeloidea and without valid taxonomic categorization (Švácha & Lawrence, 2014).

The Disteniidae collection of the Museu de Zoologia da Universidade de São Paulo (MZSP) includes about 700 specimens separated into 17 genera and 147 species. These specimens came from incorporation of other older collections such as Campos Seabra, Halik, Malkin and Melzer, among others. In the last century, this collection has grown especially by material collected by Erwin, Giesbert, Hovore, Lane, Lenko, Maller, O’Brien, Plaumann, Wappes, Zikán, and especially by Martins and Santos-Silva through identification and exchange.

MATERIAL AND METHODS

The material included in this paper belongs to Coleoptera collection of the MZSP, where the project was developed.

The photographs were taken with a Canon EOS Rebel T3i DSLR camera, Canon MP E 65 mm f/2.8-1 5X macro lens, controlled by Zerene Stacker AutoMontage software.

Label data were transcribed verbatim and additional comments or explanations are placed in square brackets. A single vertical bar separates rows within each label and a double vertical bar separates individual labels.

The species are presented in alphabetical order according to the original combination of each name.

For each species only one specimen was illustrated, if the primary type (i.e., holotype or lectotype) was in the collection, then this specimen was the illustrated. When more than one paratype is listed, the specimen illustrated was marked with the figure number.

RESULTS

Abauba iani Santos Silva & Tavakilian, 2009 (Fig. 1A-C)

Abauba ianiSantos-Silva & Tavakilian, 2009a: 23, figs. 11, 16, 52; Santos-Silva & Martins, 2010: 135, figs. 212, 227; Lingafelter et al., 2014: 78, figs. 84s t; Monné, 2023: 226.

Figure 1
Disteniidae type specimens. (A-C) Abauba iani Santos-Silva & Tavakilian, 2009, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Abauba mediorufovittata Santos-Silva & Tavakilian, 2009, male paratype (D) Dorsal view (E) Ventral view (F) Original labels (G I) Abauba napoensis Santos-Silva & Tavakilian, 2009, male paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

Paratypes (3): ♀ “PARATYPE // ECUADOR, Suc. Pr. / Lumbaqui, 2 km E / 6-15 km N, 05 Sept 98 / F.T. Hovore, coll. // genus ? / n. sp. // Abauba / iani / Santos-Silva / & Tavakilian // MZSP 40948”. (Fig. 1A-C) ♀ “PARÁTIPO // ECUADOR, Napo Pr. // 2-5 km NE Ahuano / 08/16 Sept 1998 / F.T. Hovore, coll. // new // Abauba / iani / Santos-Silva / & Tavakilian // MZSP 40949”. ♂ “PARÁTIPO // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 5, / Sta. 6 Project MAXUS Lot 915. // Abauba / iani / Santos-Silva / & Tavakilian // MZSP 40947”.

Abauba mediorufovittata Santos Silva & Tavakilian, 2009 (Fig. 1D-F)

Abauba mediorufovittataSantos-Silva & Tavakilian, 2009a: 20, figs. 10, 27, 39, 46, 51; Santos-Silva & Martins, 2010: 138, figs. 203, 225; Wappes & Arias, 2016: 4; Bezark, 2018: 41, fig. 3; Monné, 2023: 226.

Paratypes (3): ♂ “PARÁTIPO // R.F. Morris / collection // BOLIVIA: Santa Cruz / Buena Vista vic. / Flora & Fauna Hotel / 22-26/X/02 /, Morris // Wappes // Abauba / mediorufovittata / Santos-Silva / & Tavakilian // MZSP 40945”. (Fig. 1D-F) ♀ “PARÁTIPO // BOLIVIA: Santa Cruz / 3.7 km SSE Buena Vista / Hotel Flora & Fauna / 430 m., 5-15 XI 2001 // M.C. Thomas, B.K. / Dozier, blacklight / trap, tropical / transition forest // Abauba / mediorufovittata / Santos-Silva / & Tavakilian // MZSP 40946”. ♀ “PARÁTIPO // BRAZIL: Rondonia, / 62 km SW. Arique- / mês, Fzda. Rancho / Grande, X 16-1993 // Collectors: / C.W. & L.B. O’Brien // Abauba / mediorufovittata / Santos-Silva / & Tavakilian // MZSP 40944”.

Abauba napoensis Santos Silva & Tavakilian, 2009 (Fig. 1G-I)

Abauba napoensisSantos-Silva & Tavakilian, 2009a: 21, fig. 53; Santos-Silva & Martins, 2010: 136, figs. 226; Lingafelter et al., 2014: 103, figs. 114k, l; Monné, 2023: 226.

Paratype: ♂ “PARÁTIPO // ECUADOR: NAPO, Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 8 Oct. 1995 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 10, / Sta. 6 Project MAXUS Lot 1266 // Abauba / napoensis / Santos-Silva / & Tavakilian // MZSP 60588”.

America larrei Santos Silva & Tavakilian, 2009 (Fig. 2A-C)

America larreiSantos-Silva & Tavakilian, 2009b: 43, figs. 61, 64; Santos-Silva & Martins, 2010: 167, fig. 247; Morvan & Roguet, 2013: 39, fig. 78; Botero & Santos-Silva, 2020a: 152; Monné, 2023: 229.

Figure 2
Disteniidae type specimens. (A-C) America larrei Santos-Silva & Tavakilian, 2009, male paratype (A) Dorsal view (B) Lateral view (C) Original labels. (D-F) America spinosa Botero & Santos-Silva, 2020, male holotype (D) Dorsal view (E) Ventral view (F) Original labels (G-I) America tuberevexa Botero & Santos-Silva, 2020, female holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

Paratype: ♂ “PARÁTIPO // Mgne. Tortue PK 36 / 13 avril 2001 / D Faure // Paracometes sp. / Disteniini // America / larrei / Santos-Silva & / Tavakilian // MZSP 60542”.

America spinosa Botero & Santos Silva, 2020 (Fig. 2D-F)

America spinosaBotero & Santos-Silva, 2020a: 148; Monné, 2023: 231.

Holotype: ♂ “HOLOTYPE / America spinosa / Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 6 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 10, / Sta. 8 Project MAXUS Lot 887 // MZSP 60548”.

Paratypes (4): ♀ “PARATYPE / America spinosa / Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 2 Oct. 1996 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 9 Project MAXUS Lot 1719 // MZSP 60549”. ♂ “PARATYPE / America spinosa / Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 10 Feb. 1996 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 10, / Sta. 10 Project MAXUS Lot 1500 // MZSP 60550”. ♀ “PARATYPE / America spinosa / Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 7 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 8, / Sta. 10 Project MAXUS Lot 909 // MZSP 60551”. ♂ “PARATYPE / America spinosa / Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 6 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 9, / Sta. 10 Project MAXUS Lot 879 // MZSP 60552”.

America tuberevexa Botero & Santos Silva, 2020 (Fig. 2G-I)

America tuberevexaBotero & Santos-Silva, 2020a: 152; Monné, 2023: 231.

Holotype: ♀ “HOLOTYPE / America tuberevexa / Botero & Santos-Silva, 2020 // ECUADOR Napo Pr. / Napo-Galeras Rd., km 2 / 10 Sept 2000 / F.T. Hovore, coll. // MZSP 60554”.

Arietocometes terryi Botero & Santos Silva, 2021 (Fig. 3A-C)

Arietocometes terryiBotero & Santos-Silva, 2021: 207; Monné, 2023: 232.

Figure 3
Disteniidae type specimens. (A-C) Arietocometes terryi Botero & Santos-Silva, 2021, female holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D F) Arietocometes virgatus Botero & Santos-Silva, 2021, female holotype (D) Dorsal view (E) Ventral view (F) Original labels (G-I) Cometes birai Hovore & Santos-Silva, 2007, female holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

Holotype: ♀ “HOLOTYPE / Arietocometes terryi / des. Botero & Santos-Silva, 2020 // Ecuador, Napo, Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 4 Oct. 1996 / 220 m. 00°39′10″S, 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest. At Trans. 10 / Sta. 10 Project MAXUS Lot 1760 // MZSP 60585”.

Arietocometes virgatus Botero & Santos Silva, 2021 (Fig. 3D-F)

Arietocometes virgatusBotero & Santos-Silva, 2021: 200; Monné, 2023: 233.

Holotype: ♀ “HOLOTYPE / Arietocometes virgatus / des. Botero & Santos-Silva, 2020 // Ecuador, Napo / Aliñhui, 06 / Dec 2005 F.T. / Hovore, coll // MZSP 60586”.

Cometes birai Hovore & Santos Silva, 2007 (Fig. 3G-I)

Cometes biraiHovore & Santos-Silva, 2007a: 122; Swift et al., 2010: 67.

Paracometes birai;Santos-Silva & Tavakilian, 2009b: 27; Monné et al., 2017: 87; Botero & Almeida, 2019: 53; Botero & Gutiérrez, 2020: 8, Monné, 2023: 256.

Holotype: ♀ “HOLOTYPE // Cometes / eximius / Bates // [front] Costa Rica / F Nevermann / 20 VIII [19]22 / [back] Hamburg farm / Reventazon / Ebene Limon // Cometes / birai / Hovore & / Santos-Silva // Paracometes / birai / (Hovore & / Santos-Silva, 2007) // MZSP 60580” Note: There are still two handwritten labels.

Current combination: Paracometes birai.

Cometes giesberti Hovore & Santos Silva, 2007 (Fig. 4A-C)

Cometes giesbertiHovore & Santos-Silva, 2007b: 76.

Figure 4
Disteniidae type specimens. (A-C) Cometes giesberti Hovore & Santos-Silva, 2007, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Cometes hovorei Santos-Silva, 2007, female paratype (D) Dorsal view (E) Ventral view (F) Original labels (G-I) Cometes mariahelenae Hovore & Santos-Silva, 2007, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

Arietocometes giesberti;Santos-Silva & Tavakilian, 2009b: 29; Botero & Santos-Silva, 2021: 209; Monné, 2023: 232.

Paratype: ♀ “PARATYPE // Panama Bocas del Toro / 10 km NE Fortuna Dam / 3400’ May 23-26 1984 / E. Giesbert coll. // Cometes / birai / Hovore & / Santos-Silva // Arietocometes / giesberti / (Hovore & / Santos-Silva, 2007) // MZSP 60581”.

Current combination: Arietocometes giesberti.

Cometes hovorei Santos Silva, 2007 (Fig. 4D-F)

Cometes hovorei Santos-Silva in Hovore & Santos-Silva, 2007b: 95, figs. 15, 22, 29.

America hovorei;Santos-Silva & Tavakilian, 2009b: 30, fig. 30; Santos-Silva & Martins, 2010: 152, fig. 234; Le Tirant & Santos-Silva, 2014b: 1; Botero & Santos-Silva, 2020a: 152; Monné, 2023: 229.

Paratype: ♀ “PARÁTIPO // Satipo / PERU / x.1940 / A. Maller // Cometes / hovorei / Hovore & / Santos-Silva // America / hovorei / (Santos-Silva, 2007) // MZSP 60541”.

Current combination: America hovorei.

Cometes mariahelenae Hovore & Santos Silva, 2007 (Fig. 4G-I)

Cometes mariahelenaeHovore & Santos-Silva, 2007b: 80, figs. 4, 19, 33, 34.

America mariahelenae;Santos-Silva & Tavakilian, 2009b: 30, fig. 31; Santos-Silva & Martins, 2010: 169, figs. 249, 251, 252; Botero & Santos-Silva, 2020a: 147; Monné, 2023: 229.

Paratypes (3): ♀ “PARÁTIPO // BRAZIL: Rondonia, / 62 km SW. Arique- / mes, Fzda. Rancho / Grande, X 8-1993 // UV & merc. vap. light / C.W. & L.B. O’Brien // Cometes / mariahelenae / Hovore & / Santos-Silva // America / hovorei / (Hovore & / Santos-Silva, 2007) // MZSP 60543”. (Fig. 4G-I) ♀ “PARÁTIPO // BRAZIL: Rondonia, / 62 km SW. Arique- / mes, Fzda. Rancho / Grande, X 8-1993 // Collectors; / C.W. & L.B. O’Brien // Cometes / mariahelenae / Hovore & / Santos-Silva // MZSP 60544”. ♀ “PARÁTIPO // BRAZIL, Rondônia, / Ariquemes, Rio Ji- / Parana, 25.x.1986 // 09°44′S - 61°52′W / J.A. Rarpel [?] / Arm. malaise // Paracometes / laetificus / (Bates) / M.H. Galileo det. 2000 // Cometes / mariahelenae / Hovore & / Santos-Silva // America / hovorei / (Hovore & / Santos-Silva, 2007) // MZSP 60545”.

Current combination: America mariahelenae.

Cometes mediovittipennis Santos Silva & Tavakilian, 2009 (Fig. 5A-C)

Cometes mediovittipennisSantos-Silva & Tavakilian, 2009a: 9, fig. 55; Santos-Silva & Martins, 2010: 118, fig. 199; Monné et al., 2017: 85; Monné, 2023: 234.

Figure 5
Disteniidae type specimens. (A-C) Cometes mediovittipennis Santos-Silva & Tavakilian, 2009, female holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Cometes melzeri Santos-Silva & Martins, 2004, male holotype (D) Dorsal view (E) Ventral view (F) Original labels (G-I) Cometes monnei Hovore & Santos-Silva, 2007, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

Holotype: ♀ “HOLOTYPE // Est. Esp. Santo / Sta. Theresa / 17.X.28 / O Conde // Cometes / carinatus / vill. // Cometes / mediovittipennis / Santos-Silva / & Tavakilian // MZSP 40941”.

Cometes melzeri Santos Silva & Martins, 2004 (Fig. 5D-F)

Cometes melzeriSantos-Silva & Martins, 2004: 148, figs. 4, 5; Hovore & Santos-Silva, 2007b: 82, figs. 28, 38, 49; Santos-Silva & Tavakilian, 2009a: 8; Santos-Silva & Martins, 2010: 119, fig. 200; Favretto et al., 2013: 49; Monné et al., 2017: 85; Monné, 2023: 234.

Holotype: ♂ “HOLÓTIPO // Cotypus / Cotypus // 4. [unintelligible]. 25 / Mar. // Cometes / zikani / Melz. // Cometes melzeri / Santos-Silva / & Martins // MZSP 40934”.

Paratype: ♂ “PARÁTIPO // Brasilien / Nova Teutonia / 27°11′ B 52°23′ L / Fritz Plaumann / X.4.1976 / 300-500 m // Cometes melzeri / Santos-Silva / & Martins // MZSP 40935”.

Cometes monnei Hovore & Santos Silva, 2007 (Fig. 5G-I)

Cometes monneiHovore & Santos-Silva, 2007b: 81, figs. 6, 25, 26, 36, 50, 51; Santos-Silva & Tavakilian, 2009a: 8; Santos-Silva & Martins, 2010: 116, figs. 180, 216; Monné, 2023: 234.

Paratype: ♀ “PARÁTIPO // LOCAL: Taguatinga - DF / DATA: 02 XI.83 / COL: J. Dalmacio // Cometes / monnei / Hovore & / Santos-Silva // MZSP 40942”.

Cometes nearnsiHovore & Santos Silva, 2007 (Fig. 6A-C)

Cometes nearnsiHovore & Santos-Silva, 2007b: 85; Swift et al., 2010: 68.

Figure 6
Disteniidae type specimens. (A-C) Cometes nearnsi Hovore & Santos-Silva, 2007, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Cometes solangeae Hovore & Santos-Silva, 2007, female paratype (D) Dorsal view (E) Ventral view (F) Original labels (G I) Cometes soledari Martins & Galileo, 2001, male holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

Arietocometes nearnsi;Santos-Silva & Tavakilian, 2009b: 29; Botero & Santos-Silva, 2021: 211; Monné, 2023: 232.

Paratypes (3): ♀ “PARATYPE // Rio San Lorenzo, 1050 m / Tierras Morenas, Z.P. / Tenorio, Prov. Guanacaste / Costa Rica, M. Segura / 28 mar a 21 abr 1992 / L N 287800, 427600 // Costa Rica / INBIO / CR1000 / 752645 // Cometes / nearnsi / Hovore & / Santos-Silva // Arietocometes / nearnsi / (Hovore & / Santos-Silva, 2007) // MZSP 60582”. (Fig. 6A-C) ♀ “PARATYPE // Rio San Lorenzo, 1050 m / Tierras Morenas, Z.P. / Tenorio, Prov. Guanacaste / Costa Rica, M. Segura / 28 mar a 21 abr 1992 / L N 287800, 427600 // Costa Rica / INBIO / CR1000 / 414893 // Cometes / nearnsi / Hovore & / Santos-Silva // Arietocometes / nearnsi / (Hovore & / Santos-Silva, 2007) // MZSP 60583” ♂ “PARATYPE // Costa Rica Prov. Guanacaste / Macizo Miravalles, Estación Cabro / Muco, 1100 m, 26 mar - 3 abr 2003 / J.D. Gutiérrez, Tp de luz / L N 299769-411243 # 73494 // INBIO0003708207 / INBIOCRI COSTA RICA // MZSP 60584”.

Current combination: Arietocometes nearnsi.

Cometes solangeae Hovore & Santos Silva, 2007 (Fig. 6D-F)

Cometes solangeaeHovore & Santos-Silva, 2007b: 89, figs. 10, 20, 35, 46.

America solangeae;Santos-Silva & Tavakilian, 2009b: 30, fig. 18; Santos-Silva & Martins, 2010: 154, fig. 235; Monné, 2023: 230.

Paratypes (2): ♀ “PARÁTIPO // COLECAO / CAMPOS SEABRA // Rio Branco / Terr. Acre Brasil / Novembro 1961 / F.M. Oliveira / Paracometes / buqueti / (Thomson) // Cometes / solangeae / Hovore & / Santos-Silva // America / solangeae / (Hovore & / Santos-Silva, 2007) // MZSP 60546”. (Fig. 6D-F) ♀ “PARÁTIPO // BRAZIL / Benj. Constant / Rio Javary / alto amazonas / XI 1962 / Dirings // Cometes / solangeae / Hovore & / Santos-Silva // America / solangeae / (Hovore & / Santos-Silva, 2007) // MZSP 60547”.

Current combination: America solangeae.

Cometes soledariMartins & Galileo, 2001 (Fig. 6G-I)

Cometes soledariMartins & Galileo, 2001: 16, fig. 1; Santos-Silva & Martins, 2004: 147; Wappes et al., 2006: 42; Monné et al., 2017: 86.

Cupecuara soledari;Santos-Silva & Tavakilian, 2009a: 16; Santos-Silva & Martins, 2010: 130, fig. 221; Lingafelter et al., 2017: 3, fig. 1; Botero & Santos-Silva, 2020b: 648, figs. 39 42; Monné, 2023: 235.

Holotype: ♂ “HOLÓTIPO // BOLIVIA: Beni / Uyapi: Guanay / X XI 1992 // Cometes / soledari sp. n. / Holótipo ♂ / U.R. Martins & / M.H. Galileo det. 2000 // Cupecuara / soledari / (Martins & / Galileo, 2001) // MZSP 60578”.

Current combination: Cupecuara soledari.

Cometes solisi Hovore & Santos Silva, 2007 (Fig. 7A-C)

Cometes solisiHovore & Santos-Silva, 2007b: 92, figs. 12, 39, 44; Hubweber, 2008: 255.

Figure 7
Disteniidae type specimens. (A-C) Cometes solisi Hovore & Santos-Silva, 2007, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Cometes thomasi Hovore & Santos-Silva, 2007, female paratype (D) Dorsal view (E) Lateral view (F) Original labels (G-I) Cometes turnbowi Hovore & Santos-Silva, 2007, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 2 mm.

America solisi;Santos-Silva & Tavakilian, 2009b: 30; Swift et al., 2010: 68; Monné, 2023: 230.

Paratype: ♀ “PARATYPE // COSTA RICA Prov. Puntarenas R.V. / S. Golfito, estación Naranjales, 50 - / 200 m, 27 abr 2004. W. Porras, Libre, / L_S_289900_553450 # 76843 // INB0003837565 / INBIOCRI COSTA RICA // Cometes / solisi / Hovore & / Santos-Silva // America / solisi / (Hovore & / Santos-Silva, 2007) // MZSP 60553”.

Current combination: America solisi.

Cometes thomasi Hovore & Santos Silva, 2007 (Fig. 7D-F)

Cometes thomasiHovore & Santos-Silva, 2007b: 293, figs. 13, 21, 30, 31.

America thomasi;Santos-Silva & Tavakilian, 2009b: 30, fig. 32; Santos-Silva & Martins, 2010: 166, fig. 246; Le Tirant & Santos-Silva, 2014b: 1; Botero & Almeida, 2019: 54; Botero & Santos-Silva, 2020a: 152; Monné, 2023: 231.

Paratypes: ♀ “PARATIPO // Coleção / J.M. BOSQ // Peru / Satipo / XI 1942 // Paracometes / peruvianus / Villiers, 1958 // Cometes / thomasi / Hovore & / Santos-Silva // America / thomasi / (Hovore & / Santos-Silva, 2007) // MZSP 60555”. (Fig. 7D-F) ♀ “PARATIPO // Satipo / PERU / VIII / A. Maller // U. Martins det. 199 / Paracometes / peruvianus / ??? // Cometes / thomasi / Hovore & / Santos-Silva // America / thomasi / (Hovore & / Santos-Silva, 2007) // MZSP 60556”.

Current combination: America thomasi.

Cometes turnbowi Hovore & Santos Silva, 2007 (Fig. 7G-I)

Cometes turnbowiHovore & Santos-Silva, 2007b: 94, figs. 14, 41; Lingafelter et al., 2014: 337, figs. 175i, j.

Cupecuara turnbowi;Santos-Silva & Tavakilian, 2009a: 16; Lagos & Barrios, 2014: 16; Botero & Santos-Silva, 2020b: 643, figs. 10-17; Monné, 2023: 235.

Paratype: ♀ “PARATYPE // PANAMA: C.Z. / BCI, 9°9′N, 79°51′W / 980520-980603 / Pickering & Windsor // malaise trap / Cometes / turnbowi / Hovore & / Santos-Silva // Cupecuara / turnbowi / (Hovore & / Santos-Silva, 2007) // MZSP 60579”.

Current combination: Cupecuara turnbowi.

Cometes wappesi Santos Silva & Martins, 2004 (Fig. 8A-C)

Cometes wappesiSantos-Silva & Martins, 2004: 150; Monné & Hovore, 2006: 301.

Figure 8
Disteniidae type specimens. (A-C) Cometes wappesi Santos-Silva & Martins, 2004, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D F) Cometes zikani Melzer, 1926, female lectotype (D) Dorsal view (E) Ventral view (F) Original labels (G-I) Cupecuara erwini Botero & Santos-Silva, 2020, male holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Arietocometes wappesi;Santos-Silva & Tavakilian, 2009b: Lingafelter et al., 2014: 346; Botero & Santos-Silva, 2021: 213; Monné, 2023: 233.

Paratype: ♀ “PARÁTIPO // Panama Chiriqui / Cont’l Divide Trail / 12-14 May 1999 / Wappes & Morris // Cometes / wappesi / Santos-Silva / & Martins // Arietocometes / wappesi / (Santos-Silva & / Martins, 2004) // MZSP 60587”.

Current combination: Arietocometes wappesi.

Cometes zikaniMelzer, 1926 (Fig. 8D-F)

Cometes zikaniMelzer, 1926: 6; Zikán & Zikán, 1944: 4; Zikán & Wygodzinsky, 1948: 32; Villiers, 1958: 413; Zajciw, 1972: 48; Santos-Silva & Martins, 2004: 147, fig. 3; Hovore & Santos-Silva, 2007b: 82, figs. 27, 37, 48; Santos-Silva & Tavakilian, 2009a: 8; Santos-Silva & Martins, 2010: 122, fig. 217; Monné et al., 2017: 86; Monné, 2023: 234.

Lectotype: ♀ “LECTÓTIPO // Typus // Typus // 3. Virginia XII / S. Minas Ger. / Faz. Campos / 1500 m. 1915 / J. Zikán // Cometes / zikani / Melz. // MZSP 40936” (Fig. 8D-F).

Paralectotypes (3): ♀ “PARALECTÓTIPO // Cotypus // Cotypus // J.F. Zikán // [FRONT] Mimas (Brazil) / Passa Quatro / Faz. Campos / XII.17 / [VERSE] 25. XII. 1917 / Zikan // Cometes / zikani / Melz. // MZSP 40938”. ♀ “PARALECTÓTIPO // Cotypus // Cotypus // J. Zikán // IEEA // [FRONT] Virginia / S. Minas Ger. / Faz. Campos / 1500 m. / [VERSE] 21. XI. 19 // M fot. / II.71 // Cometes / zikani / Melz. // MZSP 40939”. ♀ “PARALECTÓTIPO // Cotypus // Cotypus // J.F. Zikán // [FRONT] Minas (BRAZIL) / Passa Quatro / FAZ. DOS CAMPOS / 23.XI.15 / [VERSE] 23.XI.1915 // Cometes / zikani / Melz. // MZSP 40937”.

Cupecuara erwini Botero & Santos Silva, 2020 (Fig. 8G-I)

Cupecuara erwiniBotero & Santos-Silva, 2020b: 643, figs. 18-37; Monné, 2023: 234.

Holotype: ♂ “HOLOTYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 7 Project MAXUS Lot 926 // MZSP 60566”.

Paratypes (11): ♀ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 3 Project MAXUS Lot 922 // MZSP 60577”. ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 6 Oct. 1995 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 10 Project MAXUS Lot 1230 // MZSP 60568”. ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 8 Oct. 1995 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 10, / Sta. 10 Project MAXUS Lot 1270 // MZSP 60569”. ♀ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 10 Project MAXUS Lot 929 // MZSP 60567”. ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 8 Oct. 1995 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 10, / Sta. 6 Project MAXUS Lot 1266 // MZSP 60571”. ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 3 Project MAXUS Lot 922 // MZSP 60570” ♀ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 5, / Sta. 8 Project MAXUS Lot 917 // MZSP 60572” ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 2 Oct. 1996 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 3 Project MAXUS Lot 1713 // MZSP 60574”. ♀ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 4 Oct. 1996 / 220 m 00°39′10″S 076°26′W / T.L. Erwin et al. collectors // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 10, / Sta. 3 Project MAXUS Lot 1753 // MZSP 60575”. ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 3 Project MAXUS Lot 928 // MZSP 60573”. ♂ “PARATYPE / Cupecuara erwini / des. Botero & Santos-Silva, 2020 // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 6, / Sta. 3 Project MAXUS Lot 922 // MZSP 60576”.

Distenia (Distenia) charynae Santos Silva & Hovore, 2007 (Fig. 9A-C)

Distenia (Distenia) charynaeSantos-Silva & Hovore, 2007b: 24, figs. 28, 39; Santos-Silva & Martins, 2010: 226, fig. 298; Monné, et al., 2017: 86; Botero & Almeida, 2019: 47; Monné, 2023: 236.

Figure 9
Disteniidae type specimens. (A-C) Distenia (Distenia) charynae Santos-Silva & Hovore, 2007, male holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D F) Distenia (Distenia) marcelae Santos-Silva & Hovore, 2007, male holotype (D) Dorsal view (E) Ventral view (F) Original labels (G-I) Distenia (Distenia) marinonii Botero & Almeida, 2019, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Holotype: ♂ “HOLÓTIPO // Sta. Cruz da Serra / (5 km 5 W) - RO / 4-10.XI.985 / MZ - Polo Noroeste // D. (Distenia) / charynae / Santos-Silva / & Hovore // MZSP 40629”.

Paratypes (2): ♀ “PARÁTIPO // R.F. Morris / collection // BOLIVIA: Santa Cruz / Buena Vista, F&F / Hotel, 14-17/XI/03 / Morris, Nearns, / Wappes // D. (Distenia) / charynae / Santos-Silva / & Hovore // MZSP 40631”. ♂ “PARÁTIPO // BOLIVIA, S. Cruz 4-6 K / SSE B. Vista F & F Hotel / 13-16 Nov 2003 J. / Wappes, Morris & Nearns // D. (Distenia) / charynae / Santos-Silva / & Hovore // MZSP 40630”.

Distenia (Distenia) marcelae Santos Silva & Hovore, 2007 (Fig. 9D-F)

Distenia (Distenia) marcelaeSantos-Silva & Hovore, 2007b: 19, fig. 25, 36; Santos-Silva & Martins, 2010: 231, fig. 302; Santos-Silva & Galileo, 2011: 45; Giuglaris, 2012: 61; Morvan & Roguet, 2013: 39, fig. 79; Monné et al., 2017: 86; Botero & Almeida, 2019: 55; Monné, 2023: 237.

Holotype: ♂ “HOLÓTIPO // BRASIL / ITAITUBA / (Rio Tapajos) / Estado Pará / DIRINGS // Distenia / sp. / M.A. Monné, det. 1973 // D. (Distenia) / marcelae / Santos-Silva / & Hovore // MZSP 40632”.

Distenia marinoniiBotero & Almeida, 2019 (Fig. 9G-I)

Paratypes (2): ♂ “PARATYPE / Distenia (D.) marinonii / sp. nov. / Des: Botero J.P. & Almeida, L.M. 2018 // Linhares, E.S. / X.1972 M[acho] / B. Silva // DZUP 469294 // MZSP 60557”. ♀ “Paratype / Distenia (D.) marinonii / sp. nov. / Des: Botero J.P. & Almeida, L.M. 2018 // Parque Sooretama F[emea] / Linhares / E. Santo / Brasil XI.1962 / F.M. Oliveira // DZUP 469276 // Coleção Campos Seabra // MZSP 60558” (Fig. 9G-I).

Distenia (Distenia) mermudesi Santos Silva & Hovore, 2007 (Fig. 10A-C)

Distenia (Distenia) mermudesiSantos-Silva & Hovore, 2007b: 16, figs. 2, 12, 24, 38; Santos-Silva & Martins, 2010: 223, fig. 291, 296; Monné et al., 2017a: 86; Moura & Von Groll, 2017: 461; Botero & Almeida, 2019: 47; Monné, 2023: 238.

Figure 10
Disteniidae type specimens. (A-C) Distenia (Distenia) mermudesi Santos-Silva & Hovore, 2007, male holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Distenia (Distenia) peninsularis Santos-Silva & Hovore, 2008, female paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Distenia pulchra Melzer, 1926, male lectotype (D) Dorsal view (E) Ventral view (F) Original labels. Scale bar: 1 mm.

Holotype: ♂ “HOLÓTIPO // Brasil Pará / Serra Norte / 3 ALFA / 16 X 1984 // Brasil Pará / W Franca // Distenia / agroides / Bates, 1870 / M.H. Galileo det. 2002 // Distenia (D.) / mermudesi / Santos-Silva / & Hovore // MZSP 40635”.

Paratypes (4): ♀ “PARÁTIPO // Utiariti / Rio Papagaio, Mt / 22-31.X.1966 / Lenko & Pereira // Distenia (D.) / mermudesi / Santos-Silva / & Hovore // MZSP 40636”. ♂ “PARÁTIPO // BOLIVIA Santa Cruz / 4-6k SSE Buena Vista / F&F Hotel 23-26 Oct / 2000 Wappes & Morris // D. (Distenia) / mermudesi / Santos-Silva / & Hovore // MZSP 40641”. ♀ “PARÁTIPO // R.F. Morris / collection // BOLIVIA: Santa Cruz / Buena Vista vic. / Flora & Fauna Hotel / 22-26/X/02, Morris / Wappes // D. (Distenia) / mermudesi / Santos-Silva / & Hovore // MZSP 40640”. ♂ “PARÁTIPO // BOLIVIA Santa Cruz / 4-6k SSE Buena Vista / F&F Hotel 23-26 Oct. / 2000 Wappes & Morris // D. (Distenia) / mermudesi / Santos-Silva / & Hovore // MZSP 40639”. ♂ “PARÁTIPO / BOLIVIA: Beni / Uyapi: Guanay / X XI 1992 // Distenia / agroides / Bates, 1870 / M.H. Galileo det. 2002 / Distenia (D.) / mermudesi / Santos-Silva / & Hovore // MZSP 40637”.

Distenia (Distenia) peninsularis Santos Silva & Hovore, 2008 (Fig. 10D-F)

Distenia (Distenia) peninsularisSantos-Silva & Hovore, 2008c: 14, figs. 8, 14; Botero & Almeida, 2019: 47; Monné, 2023: 238.

Paratype: ♀ “PARÁTIPO // Rancho Quemado, Pen. / Osa. Prov. Punt, COSTA / RICA, F. Quesada, Abr / 1991, L S 292500, 511000 // COSTA RICA / INBIO / CR1000 / 658662 // Distenia / (Distenia) / peninsularis / Santos-Silva & Hovore // MZSP 40679”.

Distenia pulchraMelzer, 1926 (Fig. 10G-I)

Distenia pulchraMelzer, 1926: 5; Zikán & Zikán, 1944: 4; Blackwelder, 1946: 558; Zikán & Wigodzinsky, 1948: 33; Villiers, 1959: 58.

Oculipetilus pulcher;Santos-Silva & Hovore, 2007c: 16, figs. 79, 90; Santos-Silva & Martins, 2010: 181, fig. 262; Monné et al., 2017a: 87; Monné, 2023: 256.

Distenia mirabilisVilliers, 1959: 62, figs. 7-9.

Lectotype: ♂ “Typus // Typus // [FRONT] Virginia / S. Minas Ger. / Faz. Campos / 1500 m. / J. Zikan / [VERSE] 2. XII. 1920 // Distenia / pulchra / Melz. // MZSP 40838” (Fig. 10G-I).

Paralectotypes (2): ♀ “Cotypus // Cotypus // IEEA // [FRONT] Virginia / S. Minas Ger. / Faz. Campos / 1500 m. / J. Zikan / [VERSE] 5. I. 1919 // Distenia / pulchra / Melz. // MZSP 40840”. ♀ “Cotypus // Cotypus // J.F. ZIKÁN // [FRONT] MINAS (BRAZIL) / PASSA QUATRO / FAZ. DOS CAMPOS / [VERSE] 1 jan. 1918 / Zikán // Distenia / pulchra / Melz. // MZSP 40839”.

Current combination: Oculipetilus pulcher.

Distenia (Distenia) solangeae Santos Silva & Hovore, 2007 (Fig. 11A-C)

Distenia (Distenia) solangeaeSantos-Silva & Hovore, 2007b: 21, figs. 3, 9, 26, 31; Santos-Silva & Martins, 2010: 220, figs. 289, 292, 294; Botero & Almeida, 2019: 45; Monné, 2023: 238.

Figure 11
Disteniidae type specimens. (A-C) Distenia (Distenia) solangeae Santos-Silva & Hovore, 2007, male paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D F) Distenia (Distenia) tavakiliani Santos-Silva & Hovore, 2007, male paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) bispinosa Santos-Silva & Hovore, 2008, male paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratypes (2): ♂ “PARÁTIPO // COLECAO / CAMPOS SEABRA // SINOP 12°31′S, 55°37′W / BR 163 km 500 a 600 / Mato Grosso, BRASIL / 350 m X.1974 / Alvarenga & Roppa col. // D. (Distenia) / solangeae / Santos-Silva / & Hovore // MZSP 40647”. ♂ “PARÁTIPO // COLECAO / CAMPOS SEABRA // SINOP 12°31′S, 55°37′W / BR 163 km 500 a 600 / Mato Grosso, BRASIL / 350 m X.1974 / Alvarenga & Roppa col. // D. (Distenia) / solangeae / Santos-Silva / & Hovore // MZSP 40646” (Fig. 11A-C).

Distenia (Distenia) tavakiliani Santos Silva & Hovore, 2007 (Fig. 11D-F)

Distenia (Distenia) tavakilianiSantos-Silva & Hovore, 2007b: 22, figs. 27, 39; Santos-Silva & Martins, 2010: 234, fig. 304; Botero & Almeida, 2019: 47; Monné, 2023: 239.

Paratype: ♂ “PARÁTIPO // IavH / Ei 132 / ii.2003 // COLOMBIA Amazonas / PNN Amacayacu Matamata / m2-3°41′S 70°15′W 150 m / Malaise 9-30.vii.2001 / D. Chota Leg. M.2030. // D. (Distenia) / tavakiliani / Santos-Silva & Hovore // MZSP 40648”.

Elytrimitatrix (Grossifemora) bispinosa Santos Silva & Hovore, 2008 (Fig. 11G-I)

Elytrimitatrix (Grossifemora) bispinosaSantos-Silva & Hovore, 2008b: 2; Heffern & Santos-Silva, 2016: 573; Monné, 2023: 242.

Paratypes (2): ♀ “PARATYPE // Mexico / Queretaro / 19.0 mi. W. Xilitla / 50001 / 12.VI.1983 / R. Anderson / dry oak forest // Elytrimitatrix / (Grossifemora) / bispinosa / Santos-Silva & Hovore // MZSP 40713”. ♂ “PARATYPE // Mexico / Hidalgo / 73.9 km SW Tamazunchale / 5600’ / 4 June 1987 / R. Turnbow // Elytrimitatrix / (Grossifemora) / bispinosa / Santos-Silva & Hovore // MZSP 40714” (Fig. 11G-I).

Elytrimitatrix (Grossifemora) castaneaSantos Silva & Hovore, 2008 (Fig. 12A-C)

Elytrimitatrix (Grossifemora) castaneaSantos-Silva & Hovore, 2008b: 10; Lingafelter et al., 2014: 38; Monné, 2023: 243.

Figure 12
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) castanea Santos-Silva & Hovore, 2008, male paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) charpentierae Le Tirant & Santos-Silva, 2014, male holotype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) cinnamea Santos-Silva & Hovore, 2008, female holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratypes (3): ♂ “PARÁTIPO // Panama: Panama / Llano Carti Rd., at / km 9, V 18-1993 / Coll. E.G. Riley // Distenia sp. / (near rufa Vill.) / det. E. Giesbert // Elytrimitatrix / (Grossifemora) / castanea / Santos-Silva & Hovore // MZSP 40716”. (Fig. 12A-C) ♂ “PARATYPE // 12 km N El Llano / Pan. Prov., Panama / 23/24 Jan 1993 / F.T. Hovore, coll. // Elytrimitatrix / (Grossifemora) / castanea / Santos-Silva & Hovore // MZSP 40718”. ♀ “PARATYPE // Panama: Panama Prov. / 9 km N El Llano / V 21-1993, blacklight / R. Andrews & A. Gilbert // Elytrimitatrix / (Grossifemora) / castanea / Santos-Silva & Hovore // MZSP 40717”.

Elytrimitatrix (Grossifemora) charpentierae Le Tirant & Santos Silva, 2014 (Fig. 12D-F)

Elytrimitatrix (Grossifemora) charpentieraeLe Tirant & Santos-Silva, 2014b: 2; Monné et al., 2017: 86; Monné, 2023: 243.

Holotype: ♂ “HOLOTYPE // Peru - Pasco / Pozuzo / IV.2014 / local collector // Elytrimitatrix / (Grossifemora) / charpentierae / Le Tirant & Santos-Silva // MZSP 40719”.

Paratypes (2): ♀ “PARATYPE // Peru - Pasco / Pozuzo / IV.2014 / local collector // Elytrimitatrix / (Grossifemora) / charpentierae / Le Tirant & Santos-Silva // MZSP 40721”. ♂ “PARATYPE // Peru - Pasco / Pozuzo / IV.2014 / local collector // Elytrimitatrix / (Grossifemora) / charpentierae / Le Tirant & Santos-Silva // MZSP 40720”.

Elytrimitatrix (Grossifemora) cinnamea Santos Silva & Hovore, 2008 (Fig. 12G-I)

Elytrimitatrix (Grossifemora) cinnameaSantos-Silva & Hovore, 2008c: 5; Monné et al., 2017: 86; Monné, 2023: 243.

Holotype: ♀ “HOLÓTIPO // Costa Rica, Puntar. / Golfo Dulce, 24 km W. / Piedras Brancas, 200 m, II 1992, Hanson // Elytrimitatrix / (Grossifemora) / cinnamea / Santos-Silva & Hovore // MZSP 40724”.

Elytrimitatrix (Grossifemora) clavata Santos Silva & Hovore, 2008 (Fig. 13AC)

Elytrimitatrix (Grossifemora) clavataSantos-Silva & Hovore, 2008b: 5; Santos-Silva & Hovore, 2008c: 10; Lingafelter et al., 2014: 42; Monné, 2023: 243.

Figure 13
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) clavata Santos-Silva & Hovore, 2008, male paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) cunninghami Heffern & Santos-Silva, 2016, female paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) curoei Santos-Silva & Le Tirant, 2016, male holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratypes (9): ♂ “PARÁTIPO // Catemaco / Ver. - Mexico / VI.1954 / Kissinger // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40725”. ♂ “PARÁTIPO // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40726”. (Fig. 13A-C) ♂ “PARATYPE // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40729”. ♂ “PARATYPE // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40731”. ♀ “PARATYPE // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40732”. ♂ “PARÁTIPO // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 60559”. ♂ “PARÁTIPO // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40727”. ♀ “PARÁTIPO // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40728”. ♂ “PARATYPE // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Elytrimitatrix / (Grossifemora) / clavata / Santos-Silva & Hovore // MZSP 40730”.

Elytrimitatrix (Grossifemora) cunninghami Heffern & Santos Silva, 2016 (Fig. 13D-F)

Elytrimitatrix (Grossifemora) cunninghamiHeffern & Santos-Silva, 2016: 575; Botero & Santos-Silva, 2020a: 157; Monné, 2023: 244.

Paratypes (2): ♂ “PARATYPE // Mexico, Jalisco // La Primavera 1600 m / VII 15-1998 / G. Nogueira col. // MZSP 40734”. ♀ “PARATYPE // Mexico, Jalisco / San Juan de la / Montaña 1600 m / VII 8-2010 / G. Nogueira coll. // Elytrimitatrix / (Grossifemora) / cunninghami / Heffern & Santos-Silva // MZSP 40733” (Fig. 13D-F).

Elytrimitatrix (Grossifemora) curoei Santos Silva & Le Tirant, 2016 (Fig. 13G-I)

Elytrimitatrix (Grossifemora) curoeiSantos-Silva & Le Tirant, 2016: 301; Monné et al., 2017: 86; Monné, 2023: 244.

Holotype: ♂ “HOLOTYPE // Mex.: Oaxaca / Santiago Xiacui / El Quelite 2350 m // N 17°17′33″ / O 96°23′ / 21 VI 06 D. Curoe / A. Ramirez col. // Elytrimitatrix / (Grossifemora) / curoei / Santos-Silva & Le Tirant // MZSP 40735”.

Paratype: ♀ “PARATYPE // Mexico, Oaxaca / Sierra de Juárez / 1650 m V 10/11 1997 / G. Nogueira col. // Elytrimitatrix / (Grossifemora) / curoei / Santos-Silva & Le Tirant // MZSP 40736”.

Elytrimitatrix (Grossifemora) franki Botero & Santos Silva, 2020 (Fig. 14A-C)

Elytrimitatrix (Grossifemora) frankiBotero & Santos-Silva, 2020a: 157; Monné, 2023: 244.

Figure 14
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) franki Botero & Santos-Silva, 2020, female holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) giesberti Santos-Silva & Hovore, 2008, female holotype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) guatemalana Santos-Silva & Hovore, 2008, male paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Holotype: ♀ “HOLOTYPE / Elytrimitatrix (G.) franki / Botero & Santos-Silva, 2020 // Ecuador, Loja Prov. / 18.5 km N Gonzanama / 04°08′08.5″S, 79°23′36.4″W / 22 February 2006 / F.T. Hovore, I. Swift coll // MZSP 60560”.

Elytrimitatrix (Grossifemora) giesberti Santos Silva & Hovore, 2008 (Fig. 14D-F)

Elytrimitatrix (Grossifemora) giesbertiSantos-Silva & Hovore, 2008a: 12; Santos-Silva & Hovore, 2008b: 22; Monné et al., 2017: 86; Monné, 2023: 244.

Holotype: ♀ “HOLÓTIPO // Mexcala / Gro. Mex. / VI 29.51 // H.E. Evans / Collector // Distenia / hogei / Bates // Elytrimitatrix / (Grossifemora) / giesberti / Santos-Silva & Hovore // MZSP 40737”.

Paratype: ♂ “PARÁTIPO // Mexcala / Gro. Mex. / VI 29.51 // H.E. Evans / Collector // Distenia / hogei / Bates // Elytrimitatrix / (Grossifemora) / giesberti / Santos-Silva & Hovore // MZSP 40738”.

Elytrimitatrix (Grossifemora) guatemalana Santos Silva & Hovore, 2008 (Fig. 14G-I)

Elytrimitatrix (Grossifemora) guatemalanaSantos-Silva & Hovore, 2008b: 13; Santos-Silva & Le Tirant, 2016: 302; Monné, 2023: 245.

Paratypes (2): ♂ “PARATYPE // Honduras: Copan / 6 km NW San Agustin / 19 May 2002 / R. Turnbow // Distenia / agriloides Bates / det. R. Turnbow // Elytrimitatrix / (Grossifemora) / guatemalana / Santos-Silva & Hovore // MZSP 40742”. (Fig. 14G-I) ♂ “PARATYPE // Guatemala, Zacapa / 6 km S La Unión, 4600″ / May 5-11, 1991 / E. Giesbert coll. // Elytrimitatrix / (Grossifemora) / guatemalana / Santos-Silva & Hovore // MZSP 40743”.

Elytrimitatrix (Grossifemora) hefferni Santos Silva & Hovore, 2008 (Fig. 15A-C)

Elytrimitatrix (Grossifemora) hefferniSantos-Silva & Hovore, 2008c: 4; Monné, 2023: 245.

Figure 15
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) hefferni Santos-Silva & Hovore, 2008, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) hondurenha Santos-Silva & Hovore, 2008, male paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) incognita Santos-Silva & Hovore, 2008, female holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratype: ♂ “PARATYPE // Mexico, Sonora / 5 m. E. Alamos / Aug. II 1973 / K. Stephan leg. // Elytrimitatrix / (Grossifemora) / hefferni // Santos-Silva & Hovore // MZSP 40744”.

Elytrimitatrix (Grossifemora) hondurenha Santos Silva & Hovore, 2008 (Fig. 15D-F)

Elytrimitatrix (Grossifemora) hondurenhaSantos-Silva & Hovore, 2008b: 7; Lingafelter et al., 2014: 76; Santos-Silva & Le Tirant, 2016: 302; Monné, 2023: 245.

Paratypes (3): ♀ “PARATYPE // Honduras: Olancho / La Muralla / V 24 1996, F.G. / Andrews & A.J. Gilbert / keys to hoegei ~ X // Elytrimitatrix / (Grossifemora) / hondurenha / S. Silva & Hovore // MZSP 40747”. ♀ “PARÁTIPO // Honduras, Ol. / P.N. La Muralla / late July 1992 / D. Hawks coll. // Elytrimitatrix / (Grossifemora) / hondurenha / Santos-Silva & Hovore // MZSP 40746”. ♂ “PARATYPE // Honduras: Yoro / PN Pico Pijol, mv +/ bl, 23 July 2001 / R. Turnbow // Distenia / agriloides Bates / det. R. Turnbow // Elytrimitatrix / (Grossifemora) / hondurenha / Santos-Silva & Hovore // MZSP 40745” (Fig. 15D-F).

Elytrimitatrix (Grossifemora) incognita Santos Silva & Hovore, 2008 (Fig. 15G-I)

Elytrimitatrix (Grossifemora) incognitaSantos-Silva & Hovore, 2008c: 13; Monné et al., 2017: 86; Monné, 2023: 245.

Holotype: ♀ “HOLOTYPE // Mex.: Candelaria / Loxicha, Oaxaca / 5-7 85 // Elytrimitatrix / (Grossifemora) / incognita / Santos-Silva / & Hovore // MZSP 40748”.

Elytrimitatrix (Grossifemora) mexicana Santos Silva & Hovore, 2008 (Fig. 16A-C)

Elytrimitatrix (Grossifemora) mexicanaSantos-Silva & Hovore, 2008a: 17; Santos-Silva & Hovore, 2008b: 21; Heffern & Santos-Silva, 2016: 577; Botero & Almeida, 2019: 56; Monné, 2023: 246.

Figure 16
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) mexicana Santos-Silva & Hovore, 2008, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) michelii Santos-Silva & Hovore, 2008, male holotype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) nearnsi Santos-Silva & Hovore, 2008, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratype: ♀ “PARATYPE // Mex.: Oaxaca / 5 mi NW of / Huajuapan / VII 28-81 / Peter Jump // Distenia sp / near hoegei // det. E. Giesbert // Elytrimitatrix / (Grossifemora) / mexicana / Santos-Silva & Hovore // MZSP 40752”.

Elytrimitatrix (Grossifemora) michelii Santos Silva & Hovore, 2008 (Fig. 16D-F)

Elytrimitatrix (Grossifemora) micheliiSantos-Silva & Hovore, 2008c: 10; Monné et al., 2017: 86; Monné, 2023: 246.

Holotype: ♂ “HOLOTYPE // 4000’, 15 mi. S. Valle Nacional, / Oax.: Mex. V.20, / 1971 H. Howden // Distenia / pictipes Bates / det. F. Hovore, 2004 // How // Elytrimitatrix / (Grossifemora) / michelii / Santos-Silva / & Hovore // MZSP 40753”.

Paratype: ♂ “PARATYPE // 4000’, 15 mi. S. Valle Nacional, / Oax.: Mex. V.20, / 1971 H. Howden How // Elytrimitatrix / (Grossifemora) / michelii / Santos-Silva / & Hovore // MZSP 40753”.

Elytrimitatrix (Grossifemora) nearnsi Santos Silva & Hovore, 2008 (Fig. 16G-I)

Elytrimitatrix (Grossifemora) nearnsiSantos-Silva & Hovore, 2008a: 23; Santos-Silva & Hovore, 2008b: 21; Lingafelter et al., 2014: 104; Monné, 2023: 246.

Paratype: ♂ “PARÁTIPO // Mexico, Veracruz / Los Tuxtlas Bio. Sta. / UNAM May 17-22, 1983 / C.W. & L. O’Brien // Distenia / sp / det. J.E. Wappes // Elytrimitatrix / (Grossifemora) / nearnsi / Santos-Silva & Hovore // MZSP 40755”.

Elytrimitatrix (Grossifemora) pseudovittata Santos Silva & Hovore, 2008 (Fig. 17A-C)

Elytrimitatrix (Grossifemora) pseudovittataSantos-Silva & Hovore, 2008b: 16; Monné et al., 2017: 86; Monné, 2023: 247.

Figure 17
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) pseudovittata Santos-Silva & Hovore, 2008, male holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) setosa Santos-Silva & Hovore, 2008, female paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Elytrimitatrix (Grossifemora) sharonae Santos-Silva & Hovore, 2008, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Holotype: ♂ “HOLOTYPE // Guat. 5 km N Finca / Colimas, Zunilito / Such. V 6 1966 / J.M. Campbell 6000’ // U.R. Martins det. 1976 / Distenia / vittata / Bat. / Elytrimitatrix / (Grossifemora) / pseudovittata / Santos-Silva & Hovore // MZSP 40760”.

Paratype: ♀ “PARATYPE // Guatemala. San Marcos / La Fraternidad / VI 2000-1600 m / J. Monzon coll. // Elytrimitatrix / (Grossifemora) / pseudovittata / Santos-Silva & Hovore // MZSP 40761”.

Elytrimitatrix (Grossifemora) setosa Santos Silva & Hovore, 2008 (Fig. 17D-F)

Elytrimitatrix (Grossifemora) setosaSantos-Silva & Hovore, 2008b: 17; Monné, 2023: 248.

Paratypes (2): ♀ “PARATYPE // Guatemala, Zacapa / 6 km S La Unión, 4600’ / May 5-11, 1991 / E. Giesbert col. // Elytrimitatrix / (Grossifemora) / setosa / Santos-Silva & Hovore // MZSP 40762”. (Fig. 17D-F) ♂ “PARATYPE // Guatemala, Zacapa / 6 km S La Unión, 4600’ / May 5-11, 1991 / E. Giesbert col. // Elytrimitatrix / (Grossifemora) / setosa / Santos-Silva & Hovore // MZSP 40763”.

Elytrimitatrix (Grossifemora) sharonae Santos Silva & Hovore, 2008 (Fig. 17G-I)

Elytrimitatrix (Grossifemora) sharonaeSantos-Silva & Hovore, 2008a: 16; Santos-Silva & Hovore, 2008b: 21; Lingafelter et al., 2014: 319; Monné, 2023: 248.

Paratype: ♀ “Guatml. B. Verapaz / 19-24 km N Salama / May 25-31, 1989 4500’ / J.E. Wappes // Distenia sp / det. E. Giesbert // Elytrimitatrix / (Grossifemora) / sharonae / Santos-Silva & Hovore // MZSP 40764”.

Elytrimitatrix (Grossifemora) spinifemora Santos Silva & Hovore, 2008 (Fig. 18A-C)

Elytrimitatrix (Grossifemora) spinifemoraSantos-Silva & Hovore, 2008c: 7; Monné et al., 2017: 86; Monné, 2023: 248.

Figure 18
Disteniidae type specimens. (A-C) Elytrimitatrix (Grossifemora) spinifemora Santos-Silva & Hovore, 2008, male holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Elytrimitatrix (Grossifemora) violacea Santos-Silva & Hovore, 2008, female paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Hovorestenia cleideae Botero, 2020, female paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Holotype: ♂ “HOLÓTIPO // Guatemala: Guat. / Antigua, 2000 m, 20 June 1973 / Ginter Ekis // JAC // new? // Elytrimitatrix / (Grossifemora) / spinifemora / Santos-Silva & Hovore // MZSP 40771”.

Elytrimitatrix (Grossifemora) violacea Santos Silva & Hovore, 2008 (Fig. 18D-F)

Elytrimitatrix (Grossifemora) violaceaSantos-Silva & Hovore, 2008b: 4; Lingafelter et al., 2014: 344; Monné, 2023: 248.

Paratypes (4): ♀ “PARÁTIPO // Panama: Canal / Barro Colorado Is. / 9°9′N, 79°51′W / 12 VI 1996 / Pickering - Windsor / # 6006 // Paracometes / sp. # 2 / det. J. Wappes // Paracometes / sp. # 2 / det. J.E. Wappes // MZSP 60561”. ♀ “PARATYPE // Santos-Silva & Hovore // MZSP 40781”. (Fig. 18D-F) ♀ “PARATYPE // Barro Colorado I / Panama / 6 III 1981 / RB & LS Kimsey // Elytrimitatrix / (Grossifemora) / violacea / Santos-Silva & Hovore // MZSP 40782”. ♀ “PARATYPE // Panamá, C.Z. / Coco Solo Hosp. / 9°21′N, 79°51′W / 26 Apr. 72 Stockwell // Elytrimitatrix / (Grossifemora) / violacea / Santos-Silva & Hovore // MZSP 40783”.

Hovorestenia cleideaeBotero, 2020 (Fig. 18G-I)

Hovorestenia cleideaeBotero, 2020: 2, figs. 1A, B, 2-5; Monné, 2023: 249.

Paratypes (2): ♀ “PARATYPE / Hovorestenia cleideae / sp. nov. / Det Botero J.P. 2020 // COLOMBIA Cundinamarca PNN / Chingaza Bosque Palacio 4°31′N / 73°45′W 2930 m Winkler / 18-20.viii.2000 Cifuentes L. Leg. // MZSP 60562”. ♀ “PARATYPE / Hovorestenia cleideae / sp. nov. / Det Botero J.P. 2020 // COLOMBIA Caquetá Mun. San / José del Fragua Vda. La Esmeralda / 1°20′N 76°6′W 1350 m Malaise 9- / 13.ix.2000 González E. Leg. // MZSP 60563” (Fig. 18G-I).

Novantinoe birai Santos Silva & Hovore, 2007 (Fig. 19A-C)

Novantinoe biraiSantos-Silva & Hovore, 2007a: 580, figs. 1, 13, 41; Santos-Silva & Martins, 2010: 206, fig. 281; Monné et al., 2017a: 86; Botero & Almeida, 2019: 51; Botero & Gutiérrez, 2020: 4; Monné, 2023: 250.

Figure 19
Disteniidae type specimens. (A-C) Novantinoe birai Santos-Silva & Hovore, 2007, male holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Novantinoe chemsaki Santos-Silva & Hovore, 2007, male holotype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Novantinoe cotopaxiana Santos-Silva & Hovore, 2007, male paratype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Holotype: ♂ “HOLÓTIPO // IEEA // Colombia / Fusagasugá / 20.XI.1919 / P. Apolinar // Distenia / agroides / Bates // Novantinoe / birai / Santos-Silva / & Hovore // 18.768 // MZSP 40802”.

Novantinoe chemsaki Santos Silva & Hovore, 2007 (Fig. 19D-F)

Novantinoe chemsakiSantos-Silva & Hovore, 2007a: 588, figs. 26, 52; Santos-Silva & Martins, 2010: 196, fig. 272; Monné et al., 2017a: 86; Botero & Almeida, 2019: 50; Monné, 2023: 250.

Holotype: ♂ “HOLÓTIPO // Colombia, Magdal. / 820 m. 20 mi. E. / Santa Marta / VI.30.73 J. Helava // U. Martins det. 1998 / Distenia / batesi ? / Vill. // Novantinoe / chemsaki / Santos-Silva / & Hovore // MZSP 40807”.

Novantinoe cotopaxiana Santos Silva & Hovore, 2007 (Fig. 19G-I)

Novantinoe cotopaxianaSantos-Silva & Hovore, 2007a: 594, figs. 30, 48; Santos-Silva & Martins, 2010: 195, fig. 271; Botero & Almeida, 2019: 50; Monné, 2023: 250.

Paratypes (4): ♀ “PARATYPE // Ecuador, Cotopaxi Pr., / vic. Las Pampas, / 11/12 April 1997 / F.T. Hovore, coll. // Novantinoe / cotopaxiana / Santos-Silva / & Hovore // MZSP 40813”. ♂ “PARATYPE // Ecuador, Cotopaxi Pr., / vic. Las Pampas, / 11/12 April 1997 / F.T. Hovore, coll. // Novantinoe / cotopaxiana / Santos-Silva / & Hovore // MZSP 40812”. ♀ “PARATYPE // Ecuador, Cotopaxi Pr. / 2 km N Las Pampas, 1450 m / April 10-12 1997 / E. Giesbert, F. Hovore // Novantinoe / cotopaxiana / Santos-Silva / & Hovore // MZSP 40811”. ♂ “PARATYPE // Ecuador, Cotopaxi Pr., / vic. Las Pampas, / 11/12 April 1997 / F.T. Hovore, coll. // n. sp. Keys to / mathans // Novantinoe / cotopaxiana / Santos-Silva / & Hovore // MZSP 40810” (Fig. 19G-I).

Novantinoe cristinae Santos Silva & Hovore, 2007 (Fig. 20A-C)

Novantinoe cristinaeSantos-Silva & Hovore, 2007a: 598, figs. 31, 34, 57; Santos-Silva & Le Tirant, 2016: 299, figs. 6, 12; Botero & Almeida, 2019: 50; Monné, 2023: 251.

Figure 20
Disteniidae type specimens. (A-C) Novantinoe cristinae Santos-Silva & Hovore, 2007, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Novantinoe hovorei Santos-Silva, 2007, male paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G-I) Novantinoe iani Santos-Silva & Hovore, 2007, male holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratype: ♀ “PARATYPE // GUAT. BAJA VERAPAZ / VIC. BIOTOPO DEL QUETZAL / 5200, MAY 25 - JUN 1, 1989 / E. GIESBERT, COLL. // Novantinoe / cristinae / Santos-Silva & Hovore // MZSP 40823”.

Novantinoe hovorei Santos Silva, 2007 (Fig. 20D-F)

Novantinoe hovorei Santos-Silva in Santos-Silva & Hovore, 2007a: 589, figs. 27, 49; Swift et al., 2010: 68; Lingafelter et al., 2014: 77; Botero & Almeida, 2019: 50; Heffern et al., 2019: 573, fig. 35; Monné, 2023: 252.

Paratypes (2): ♂ “PARATYPE // Sector Las Pailas, P.N. Rincón de la Vieja, / Prov. Guana, COSTA RICA. 800 m. 6-7 / Jun 1994, D. García, L N 307300_388600 / # 3050 // COSTA RICA / INBIO / CRI001 / 908317 / Novantinoe / hovorei / Santos-Silva // MZSP 40824”. (Fig. 20D-F) ♀ “PARATYPE // Sector Las Pailas, P.N. Rincón de la Vieja, / Prov. Guana, COSTA RICA. 800 m. 6-7 / Jun 1994, D. García, L N 307300_388600 / # 3050 // COSTA RICA / INBIO / CRI001 / 908316 / Novantinoe / hovorei / Santos-Silva // MZSP 40825”.

Novantinoe iani Santos Silva & Hovore, 2007 (Fig. 20G-I)

Novantinoe ianiSantos-Silva & Hovore, 2007a: 601, figs. 32, 48; Santos-Silva & Martins, 2010: 200, fig. 276; Le Tirant & Santos-Silva, 2014b; 2; Monné et al., 2017a: 86; Botero & Almeida, 2019: 56; Botero & Gutiérrez, 2020: 8; Monné, 2023: 253.

Holotype: ♂ “HOLOTYPE // PERU MADRE de DIOS / Rio Alto Madre de Dios / Pantiacolla Lodge 410-700 m / 12°39.4′S, 71°13.9′W / D. Brzoska 25 X 2000 // keys to / equatoriensis // Novantinoe / iani / Santos-Silva / & Hovore // MZSP 40830”.

Paratypes (3): ♀ “PARÁTIPO // ECUADOR, Napo Pr. 2 km / N Ahuano, malaise trap / 05-15 Sept 2000 / F.T. Hovore, coll. // n. sp. keys to / equatoriensis // Novantinoe / iani / Santos-Silva / & Hovore // MZSP 40832”. ♀ “PARÁTIPO // ECUADOR, Napo Pr. / 25 km E. Atahualpa / 10-15 Sept 1998 / F.T. Hovore, coll. // Novantinoe / iani / Santos-Silva / & Hovore // MZSP 40833”. ♀ “PARÁTIPO // ECUADOR, Napo Pr. / Jatun Sacha Reserve, / 04-06 Sept 2000 / F.T. Hovore, coll. // Novantinoe / iani / Santos-Silva / & Hovore // MZSP 40831”.

Novantinoe lezamai Santos Silva & Hovore, 2007 (Fig. 21A-C)

Novantinoe lezamaiSantos-Silva & Hovore, 2007a: 599, figs. 33, 59; Santos-Silva & Martins, 2010: 194, fig. 270; Monné et al., 2017: 87; Botero & Almeida, 2019: 50; Botero & Gutiérrez, 2020: 9; Monné, 2023: 253.

Figure 21
Disteniidae type specimens. (A-C) Novantinoe lezamai Santos-Silva & Hovore, 2007, female holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Novantinoe lingafelteri Santos-Silva & Hovore, 2007, female paratype (D) Dorsal view (E) Ventral view (F) Original labels. (G I) Novantinoe nogueraiSantos-Silva & Le Tirant, 2016, male holotype (D) Dorsal view (E) Ventral view (F) Original labels. Scale bar: 1 mm.

Holotype: ♀ “HOLOTYPE // ECUADOR, Napo Pr. / Napo-Galeras, km 1-2 / 02 Oct 1987 / F.T. Hovore, coll. // Novantinoe / lezamai / Santos-Silva / & Hovore // MZSP 40834”.

Paratype: ♀ “PARATYPE // ECUADOR, Napo Pr. / Napo-Galeras Rd., km 2 / 10 Sept 2000 / F.T. Hovore, coll. // new // Novantinoe / lezamai / Santos-Silva / & Hovore // MZSP 40835”.

Novantinoe lingafelteri Santos Silva & Hovore, 2007 (Fig. 21D-F)

Novantinoe lingafelteriSantos-Silva & Hovore, 2007a: 583, figs. 15, 45; Lingafelter et al., 2014: 90, figs. 99e, f; Botero & Almeida, 2019: 51; Monné, 2023: 253.

Paratype: ♀ “PARATYPE // PANAMA Colon Pr. / Fort Sherman / 9°17′N 79°59′W / 2 June 2001 / leg. F. Odegaard // On Dendropanax arboreus // CER 91 // MZSP 40829 // Novantinoe / lingafelteri / Santos-Silva / & Hovore // MZSP 40829”.

Novantinoe noguerai Santos Silva & Le Tirant, 2016 (Fig. 21G-I)

Novantinoe nogueraiSantos-Silva & Le Tirant, 2016: 300, figs. 8-11; Monné et al., 2017: 87; Botero & Almeida, 2019: 51; Monné, 2023: 254.

Holotype: ♀ “HOLOTYPE // MEX; CHIAPAS / EL ROSARIO 1800 M / 2 VII 95 / CUROE HOPKINS // Novantinoe / noguerai / Santos-Silva & / Le Tirant // MZSP 40828”.

Novantinoe payettei Santos Silva & Le Tirant, 2016 (Fig. 22A-C)

Novantinoe payetteiSantos-Silva & Le Tirant, 2016: 298, figs. 1-5; Monné et al., 2017: 87; Botero & Almeida, 2019: 50; Monné, 2023: 254.

Figure 22
Disteniidae type specimens. (A-C) Novantinoe payettei Santos-Silva & Le Tirant, 2016, male holotype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Novantinoe rileyi Santos-Silva & Hovore, 2007, female holotype (D) Dorsal view (E) Ventral view (F) Original labels. (G-H) Novantinoe solisi Santos-Silva & Hovore, 2007, male holotype (D) Dorsal view (E) Ventral view (F) Original labels. Scale bar: 1 mm.

Holotype: ♂ “HOLOTYPE // NICARAGUA: JINO. / CERRO HORNOS / 1500M 4/VI/94 / ENCINO-PINO / LUZ HG. D. CUROE // Novantinoe / payettei / Santos-Silva & / Le Tirant // MZSP 40836”.

Novantinoe rileyi Santos Silva & Hovore, 2007 (Fig. 22D-F)

Novantinoe rileyiSantos-Silva & Hovore, 2007a: 576, figs. 29, 71; Santos-Silva & Martins, 2010: 209, fig. 283; Monné et al., 2017: 87; Botero & Almeida, 2019: 51; Botero & Gutiérrez, 2020: 9; Monné, 2023: 254.

Holotype: ♀ “HOLÓTIPO // ECUADOR, Napo Pr. / Jatun Sacha Res. HQ / 27-30 Sept 1997 / F.T. Hovore, coll. // n. sp. keys to / [unintelligible] // Novantinoe / rileyi / Santos-Silva / & Hovore // MZSP 40826”.

Paratype: ♂ “PARÁTIPO // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 5, / Sta. 5 Project MAXUS Lot 914 // Novantinoe / rileyi / Santos-Silva / & Hovore // MZSP 40827”.

Novantinoe solisi Santos Silva & Hovore, 2007 (Fig. 22G-I)

Novantinoe solisiSantos-Silva & Hovore, 2007a: 585, figs. 5, 20, 54; Swift et al., 2010: 68; Monné et al., 2017: 87; Botero & Almeida, 2019: 50; Monné, 2023: 255.

Holotype: ♂ “HOLÓTIPO // MZUSP / Adquirido / W. Zikán // COSTA RICA / F. NEVERMANN // TURRIALBA / 800 M / SLG. SCHILO // Distenia / ? // [unintelligible] // Novantinoe / solisi / Santos-Silva / & Hovore // MZSP 40808”.

Paratype: ♂ “PARÁTIPO // ECUADOR: NAPO Res. Ethnica / Waorani, 1 km S. Onkone Gare / Camp, Trans. Ent. 9 Oct. 1994 / 220 m 00°39′10″S 076°26′W / T.L. Erwin, et. al. // Insecticidal fogging of mostly bare / green leaves, some with covering / of lichenous or bryophytic plants in / terre firme forest At Trans 5, / Sta. 5 Project MAXUS Lot 914 // Novantinoe / rileyi / Santos-Silva / & Hovore // MZSP 40827”.

Novantinoe thomasi Santos Silva & Hovore, 2007 (Fig. 23A-C)

Novantinoe thomasiSantos-Silva & Hovore, 2007a: 584, figs. 16, 44; Lingafelter et al., 2014: 333, figs. 170i j; Botero & Almeida, 2019: 50; Monné, 2023: 255.

Figure 23
Disteniidae type specimens. (A-C) Novantinoe thomasi Santos-Silva & Hovore, 2007, female paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Paracometes ericae Martins & Galileo, 1994, female holotype (D) Dorsal view (E) Ventral view (F) Original labels. (G-H) Pseudocometes harrisoni Le Tirant & Santos-Silva, 2014, male holotype (G) Dorsal view (H) Ventral view (I) Original labels. Scale bar: 1 mm.

Paratype: ♀ “PARÁTIPO // GUAT. Zacapa / 3 km S. La Unión / 1400 m. 6.VI.1991 / H & A Howden // Novantinoe / thomasi / Santos-Silva / & Hovore // MZSP 40809”.

Paracometes ericaeMartins & Galileo, 1994 (Fig. 23D-F)

Paracometes ericaeMartins & Galileo, 1994: 80, fig. 2.

Cometes ericae;Santos-Silva & Martins, 2004: 147.

Abauba ericae;Santos-Silva & Tavakilian, 2009a: 20; Santos-Silva & Martins, 2010: 139, fig. 228; Martins, Galileo & Limeira-de-Oliveira, 2011: 276; Monné et al., 2017: 85; Monné, 2023: 226.

Holotype: ♂ “HOLÓTIPO // Palmas, TO / S. do Lajeado / Fazenda Céu / 17.XI.1992 / Exp. MCN MZSP // Paracometes / ericae sp. n. / Martins & / M.H. Galileo det. 1994 // Abauba / ericae / (Martins & / Galileo, 1994) // MZSP 40943”.

Current combination: Abauba ericae.

Pseudocometes harrisoni Le Tirant & Santos Silva, 2014 (Fig. 23G-I)

Pseudocometes harrisoniLe Tirant & Santos-Silva, 2014a: 2, figs. 1-3; Le Tirant & Santos-Silva, 2014b: 2; Monné et al., 2017: 87; Monné, 2023: 260.

Holotype: ♂ “HOLOTYPE // PERU: Pasco, / Pozuzo, VIII 2013 / Local collector // Coll. S. Le Tirant / Leg. P. Demez / sp. nov. ? // Pseudocometes / harrisoni / Le Tirant & / Santos-Silva // MZSP 40880”.

Paratype: ♂ “PARÁTIPO // PERU: Pasco, / Pozuzo, VIII 2013 / Local collector // Coll. S. Le Tirant / Leg. P. Demez / sp. nov. ? // Pseudocometes / harrisoni / Le Tirant & / Santos-Silva // MZSP 40881”.

Villiersicometes absalom Tavakilian & Santos Silva, 2012 (Fig. 24A-C)

Villiersicometes absalomTavakilian & Santos-Silva, 2012: 1, 2 figs.; Dalens, 2013: 3; Morvan & Roguet, 2013: 39, fig. 80; Monné, 2023: 258.

Figure 24
Disteniidae type specimens. (A-C) Villiersicometes absalom Tavakilian & Santos-Silva, 2012, male paratype (A) Dorsal view (B) Ventral view (C) Original labels. (D-F) Villiersicometes galileoae Botero & Almeida, 2019, male paratype (D) Dorsal view (E) Ventral view (F) Original labels. Scale bar: 1 mm.

Paratype: ♂ “PARATYPE // Haute Courcibo (DZ EDF) / 16 juillet 1990 GUYANE / Piégeage lumineux / Jean-Marie BALOUP leg. // Villiersicometes / absalom / Tavakilian & / Santos-Silva // MZSP 40873”.

Villiersicometes galileoaeBotero & Almeida, 2019 (Fig. 24D-F)

Villiersicometes galileoaeBotero & Almeida, 2019: 53, figs. 33 35; Monné, 2023: 258.

Paratypes (2): ♂ “PARATYPE // Villiersicometes galileoae / sp. nov. / Des: Botero, J.P. & Almeida, L.M. 2018 // COLECAO / CAMPOS SEABRA // Parque Sooretama / LINHARES E. Santo / Brasil X.1963 / F.M. Oliveira // DZUP / 469258 // MZSP 60564”. ♂ “PARATYPE // Villiersicometes galileoae / sp. nov. / Des: Botero, J.P. & Almeida, L.M. 2018 // COLECAO / CAMPOS SEABRA // Parque Sooretama / LINHARES E. Santo / Brasil X.1963 / F.M. Oliveira // DZUP / 469260 // MZSP 60565” (Fig. 24D-F).

  • FUNDING INFORMATION:
    JPB: post-doctoral fellow from Fundação de Amparo à Pesquisa do Estado de São Paulo” (FAPESP) (process number 2017/178980).
  • Published with the financial support of the “Programa de Apoio às Publicações Científicas Periódicas da Universidade de São Paulo”

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Edited by

  • Edited by:
    Simone Policena Rosa

Publication Dates

  • Publication in this collection
    22 Nov 2024
  • Date of issue
    2024

History

  • Received
    14 Jan 2024
  • Accepted
    15 Feb 2024
  • Published
    01 Mar 2024
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